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Page 1 ï~~2010 THE MICHIGAN BOTANIST THE VASCULAR FLORA OF SEBERT PROPERTY, LAPORTE COUNTY, INDIANA Scott Namestnik Keith Board JFNew 616 South Washington Street 708 Roosevelt Road Bremen, Indiana 46506 Walkerton, Indiana 46574 email@example.com firstname.lastname@example.org ABSTRACT Sebert Property is comprised of an approximately 40-acre matrix of hummocky wet to mesic flatwoods, dry sandy upland flats with seasonally inundated depressions, mesophytic forests and a buttonbush pond. A remarkable assemblage of 362 taxa of native vascular plants occurs here. Twenty (20) species are listed as endangered, threatened, rare (ETR) or extirpated in Indiana; additional taxa are uncommon in the Chicago Region and Indiana and 13 species are presented as new records for LaPorte County. Keywords: Vascular Flora, Flatwoods, LaPorte County, Sebert Property INTRODUCTION Sebert Property is a 39.2-ac (15.8-ha) parcel that was donated to the LaPorte County Parks Foundation on 1 September 2000 by the late Wilmer and Rose Sebert, becoming the fifth county park in LaPorte County. No development has occurred at the site since that time. The property consists of swamp forest, early successional forest, mesophytic forest, upland forest, buttonbush swamp, sand barrens, seasonal pond, open water/mudflat, abandoned access road and roadside/roadside ditch communities. Portions of the property have been subjected to anthropogenic disturbances in the past, including excavation, likely for sand mining, and planting of pine trees; other portions are intact natural communities. A large percentage of the site consists of a community classified as boreal flatwoods by the Indiana Department of Natural Resources and The Nature Conservancy; Jacquart et al. (2002) describe the boreal flatwoods community as being dominated by Quercus palustris, Acer rubrum and Ulmus americana, with characteristic species including Betula papyrifera and Ilex verticillata. We performed a thorough inventory of the vascular flora of Sebert Property in 2007 and 2008 because there is a lack of published data on sites with boreal flatwoods and associated communities in northern Indiana. In addition, an earlier informal study showed that Sebert Property was comprised of a rich flora that includes endangered, threatened and rare (ETR) plant species, numerous plants of high conservation value and several plant species not previously known from LaPorte County.
Page 2 ï~~THE MICHIGAN BOTANIST Vol. 49 THE MICHIGAN BOTANIST Vol. 49 FIGURE 1. Soils map. BtA = Brems fine sand, 0-3% slopes; Cd = Cheektowaga fine sandy loam; Hh = Histosols and Aquolls; Mx = Morocco loamy fine sand; Nf = Newton loamy fine sand; OaC = Oakville fine sand, 4-12% slopes; Sa = Saugatuck-Pipestone complex. Location Sebert Property is located approximately 6 mi northeast of Michigan City, Indiana, on the south side of County Road 925 North, 0.6 mi west of County Road 300 West, directly south of Wintergreen Woods Nature Preserve in northern LaPorte County. The property is 1.3 mi south of the Michigan state line in Springfield Township, in the southeast of the southwest of Section 16, Township 38 North, Range 3 West (44Â~44'N, 86046'W). Soils and Geology The property is located in the Oakville-Morocco-Brems soil unit; seven soil map units are present on the site (Figure 1) (USDA 1982). Moderately well drained to well drained fine sands (Brems fine sand; Oakville fine sand) are present in a low ridge that runs east-west across the site. Very poorly drained organic soils (Histosols and Aquolls) are present in a depression in the northwest corner of the site, and poorly to somewhat poorly drained soil (Saugatuck-Pipestone complex) exists along the north boundary of the site. The remainder of the property consists of fine sands and fine sandy loams that are somewhat to very poorly drained (Cheektowaga fine sandy loam; Morocco loamy fine sand; Newton loamy fine sand). Sand deposits at Sebert Property and in much of northwest Indiana are thought to be remnants of ancient beaches of glacial Lake Chicago, the precursor of Lake Michigan (Greenberg 2002). Natural Communities Located in the Northwestern Morainal Natural Region, Sebert Property is near the boundary between the Valparaiso Moraine Section, the Chicago Lake Plain Section and the Lake Michigan Border Section (Homoya et al. 1985). The
Page 3 ï~~2010 THE MICHIGAN BOTANIST Valparaiso Moraine Section is characterized by forest to the east and prairie to the west, as well as fen, bog, lake, marsh, savanna, seep spring and swamp communities, on generally well drained calcareous silty clay loam soils; the Chicago Lake Plain Section is characterized by marsh, lake, sand savanna, sand prairie, forest and swamp communities on mostly acidic sandy soils; the Lake Michigan Border Section is characterized by beach, dune, forest, savanna and panne communities on sand and muck soils (Homoya et al. 1985). The plant communities and soil chemistry at Sebert Property are most similar to the communities that characterize the Chicago Lake Plain Section to the west, as marsh, swamp and forest communities are present on sandy and mostly acidic soils (USDA 1982). Of the swamp community behind the dunes of Lake Michigan, Swink & Wilhelm (1994) state that "this system is one of the richest and most complicated systems in our region... characterized by a complex hydrology and... interspersed by gentle rises, shallow depressions, and hummocks.. " Sebert Property fits this description quite well. In addition, several degraded communities including old-field, sand blowout and young successional forest are present. Historically, the central early successional forest, central sand barrens and west sand barrens are presumed to have been a sand ridge. This area has been cleared of vegetation and leveled within the past 40 years. MATERIALS AND METHODS Floristic Inventory A meander survey following the methods of Goff et al. (1982) was conducted in 2005 by the authors with occasional assistance from others, resulting in a plant list of 292 taxa. For that study, collections were made for identification purposes but no attempt was made to collect voucher specimens for the entire flora. The resulting species list from 2005 was used as a guide for this more thorough study also conducted by meander survey in 2007 and 2008. The current study resulted in substantial additions to the list of known vascular plants at Sebert Property. Plant communities were generally mapped (Figure 2) as part of the study to document location of occurrence of collected specimens. With a few noted exceptions, a specimen of each taxon was collected and deposited at the herbarium of The Morton Arboretum, Lisle, Illinois (MOR). For a few conservative species that occurred in FIGURE 2. Plant communities at Sebert Property. AR = Abandoned road; BBS = Buttonbush swamp; CESF = Central early successional forest; CSB = Central sand barrens; EESF = East early successional forest; NESF..= Northeast swamp forest; NWSF = North"west swamp forest; OW/M = Open water/mudflat; RS/D = Roadside and ditch; SESEP = Southeast pond; SP = Small pond; SMF = South mesophytic forest; SEUF = M Southeast upland forest; SSF = South swamp forest; WSB = West sand barrens.
Page 4 ï~~THE MICHIGAN BOTANIST Vol. 49 very small numbers, only photographic vouchers were submitted. The primary sources for plant identification were Gleason & Cronquist (1991) and Swink & Wilhelm (1994). Additional resources consulted for the identification of plant species included Deam (1940), Fernald (1950), Flora of North America Editorial Committee, eds. (1993+), Hitchcock (1950), Holmgren (1998), Mohlenbrock (2002), Steyermark (1963), Voss (1972, 1985, 1996) and Yatskievych (1999, 2006). Nomenclature in the species list follows The PLANTS Database (USDA, NRCS 2009). Floristic Quality Assessment Floristic Quality Assessment (FQA) was used to evaluate the natural area potential of the site and to allow for comparison with other sites where inventories have been conducted. Coefficient of conservatism values (C-values) for Indiana were obtained from Rothrock (2004). The methods outlined by Swink & Wilhelm (1994) and Rothrock (2004) for determining floristic quality were followed. RESULTS Floristic Inventory A total of 434 vascular plant taxa representing 245 genera and 90 families were observed and identified to at least the species level (Appendix 1). Of these, 362 (83%) are native to the Chicago Region; seventy-two (17%) are non-native. These figures consider the following species, which are native in Indiana but thought to be non-native in the Chicago Region or intentionally introduced at the site, as non-native: Andropogon virginicus, Pinus virginiana, Robinia pseudoacacia, Thuja occidentalis and Tridens flavus var. flavus. Many of the non-native plants occur in disturbed ground on the edge of County Road 925 North along the northern site boundary. Four additional taxa were observed but not identified to species: two species of Platanthera, one species of Typha and one species of Vernonia. Several individuals of one of the species of Platanthera (SAN PH-6 [MOR]) were observed, but none produced flowers during the study. One individual each of the other species of Platanthera (SAN PH-8 [MOR]), Vernonia (no collection) and Typha (no collection) were observed, but none of these produced flowers during the study. The Typha sp. was found along the right-of-way along County Road 925 North and was mowed prior to anthesis. These taxa were not included in the total species count or the Floristic Quality Assessment. Pteridophytes were represented by 22 taxa (5%), gymnosperms were represented by six taxa (1%) and angiosperms were represented by 406 taxa (94%). Poaceae was the most common family, with 51 taxa (12%) identified. Asteraceae and Cyperaceae were also well represented, with 44 taxa (10%) each. Rosaceae was the fourth most common family, with 25 taxa (6%) collected. Carex was the most well represented genus, with 35 taxa (8%) documented. Swink & Wilhelm (1994), in their discussion of Woodwardia areolata (L.) T. Moore, note that "the dune region is a virtual repository of both boreal and coastal-plain disjuncts and relicts." There are numerous plants at Sebert Property that are more commonly encountered outside of the Chicago Region, some of which may be remnants from a time when the climate was cooler, or from when coastal areas were further inland. The term "boreal" is certainly applied differently at every latitude. In Indiana and the Chicago region, this description is used
Page 5 ï~~2010 THE MICHIGAN BOTANIST TABLE 1. Boreal relicts (Swink & Wilhelm 1994) at Sebert Property. Betula papyrifera Marsh. Betula populifolia Marsh. Coptis trifolia (L.) Salisb. Juniperus communis L. Maianthemum canadense Desf. Panax trifolius L. Pinus strobus L. Pyrola elliptica Nutt. Thuja occidentalis L. Planted at site Trientalis borealis Raf. TABLE 2. Coastal plain disjuncts (Swink & Wilhelm 1994) at Sebert Property. Bartonia virginica (L.) Britton, Sterns & Poggenb. Bidens discoidea (Torr. & A. Gray) Britton Carex albolutescens Schwein. Carex longii Mack. Carex seorsa Howe Juncus scirpoides Lam. Linum striatum Walter Lycopodiella inundata (L.) Holub Polygonum careyi Olney Rhexia virginica L. Triadenum virginicum (L.) Raf. Woodwardia virginica (L.) Sm. for northern species with limited distribution near Lake Michigan. Similarly, coastal plain disjuncts, as referred to in Indiana and the Chicago region, are those plants that are much more common on the coastal plain and outer piedmont of the Mid-Atlantic states, but that are much more restricted in distribution in the Great Lakes states. Using categorizations from Swink & Wilhelm (1994), nine naturally occurring boreal relicts (Table 1) and 12 naturally occurring coastal plain disjuncts (Table 2) were observed. Examples of boreal relicts are shown in Figures 3-7 and a coastal plain disjunct is shown in Figure 8. Many other species characteristic of the boreal flatwoods community were documented. Floristic Quality Based on the taxa identified, Sebert Property has a mean C-value of 3.7 (native mean C-value of 4.4) and a floristic quality index (FQI) of 76.0 (native FQI of 83.3). Sites with mean C-values of 4.5 or higher or FQI values of 45 are usually representative of intact natural plant communities (Swink & Wilhelm 1994). As shown in Figure 9, the C-value distribution of native species generally follows a bell curve, with the greatest percentage of native species having a C-value of 4 (63 species, 17%). Species with a C-value of 4 occasionally thrive in communities with anthropogenic disturbance (Rothrock & Homoya 2005). Despite past disturbance at Sebert Property, 31 species (9%) have a C-value of 9 or 10 (Table 3). These plants are thought to be restricted to remnant landscapes that appear to have suffered very little post-settlement trauma (Rothrock 2004).
Page 6 ï~~THE MICHIGAN BOTANIST Vol. 49 THE MICHIGAN BOTANIST Vol. 49 FIGURE 3. Boreal relict Coptis trifolia (L.) Salisb. in swamp forest, April 25, 2009. Photograph by Keith Board. FIGURE 4. Boreal relict Maianthemum canadense Desf. in swamp forest, May 26, 2008. Photograph by Keith Board.
Page 7 ï~~2010 THE MICHIGAN BOTANIST 2010 THE MICHIGAN BOTANIST FIGURE 5. Boreal relict Panax trifolius L. with Viola rostrata Pursh in mesophytic forest, May 17, 2008. Photograph by Keith Board. FIGURE 6. Boreal relict Pyrola elliptica Nutt. in swamp forest, June 28, 2008. Photograph by Keith Board.
Page 8 ï~~THE MICHIGAN BOTANIST Vol. 49 THE MICHIGAN BOTANIST Vol. 49 FIGURE 7. Boreal relict Trientalis borealis Raf. in swamp forest, May 21, 2008. Photograph by Keith Board. FIGURE 8. Coastal plain disjunct Woodwardia virginica (L.) Sm. in roadside ditch, July 16, 2009. Photograph by Keith Board.
Page 9 ï~~2010 THE MICHIGAN BOTANIST 2010 THE MICHIGAN BOTANIST 70 f 60 50 ' 40 z 0 30 20 z 10 0 1 2 3 4 5 6 7 8 9 10 C-value FIGURE 9. C-value distribution of native species at Sebert Property. TABLE 3. Species with C-value of 9 or 10 (Rothrock 2004) at Sebert Property. Betula populifolia Marsh. 10 Brachyelytrum aristosum (Michx.) Trel. 10 Carex arctata Boott ex Hook. 10 Carex brunnescens (Pers.) Poir. 10 Carex debilis Michx. var. rudgei L.H. Bailey 10 Carexfolliculata L. 10 Carex seorsa Howe 10 Coptis trifolia (L.) Salisb. 10 Dryopteris intermedia (Muhl. ex Willd.) A. Gray 10 Juniperus communis L. 10 Lonicera canadensis Bartram ex Marsh. 10 Lycopodium tristachyum Pursh 10 Polygonum careyi Olney 10 Sparganium americanum Nutt. 10 Torreyochloa pallida (Torr.) Church var. pallida 10 Triadenum virginicum (L.) Raf. 10 Trientalis borealis Raf. 10 Viola macloskeyi Lloyd ssp. pallens (Banks ex Ging) M.S. Baker 10 Woodwardia virginica (L.) Sm. 10 Alnus incana (L.) Moench ssp. rugosa (Du Roi) R.T. Clausen 9 Carex albolutescens Schwein. 9 Carex tonsa (Fernald) E.P. Bicknell var. rugosperma (Mack.) Crins 9 Carex tonsa (Fernald) E.P. Bicknell var. tonsa 9 Juncus scirpoides Lam. 9 Osmunda cinnamomea L. 9 Panax trifolius L. 9 Platanthera clavellata (Michx.) Luer 9 Salix myricoides Muhl. var. myricoides 9 Vaccinium corymbosum L. 9 Veronica scutellata L. 9 Viola lanceolata L. 9
Page 10 ï~~10 THE MICHIGAN BOTANIST Vol. 49 TABLE 4. Indiana endangered, threatened, rare and watch list species (Indiana Department of Natural Resources 2010) at Sebert Property. Extirpated Lonicera canadensis Bartram ex Marsh. Endangered Betula populifolia Marsh. Brachyelytrum aristosum (Michx.) Trel.2 Carex arctata Boott ex Hook. Carex brunnescens (Pers.) Poir. Threatened Geranium robertianum L. Juncus scirpoides Lam. Rare Carex debilis Michx. var. rudgei L.H. Bailey Carexfolliculata L. Carex seorsa Howe Juniperus communis L. Lycopodium obscurum L. Watch List Alnus incana (L.) Moench ssp. rugosa (Du Roi) R.T. Clausen Betula papyrifera Marsh. Chimaphila maculata (L.) Pursh Coptis trifolia (L.) Salisb. Huperzia lucidula (Michx.) Trevis. Linum striatum Walter Lycopodium clavatum L. Lycopodiella inundata (L.) Holub Rubus enslenii Tratt. Thuja occidentalis L.1 Polygonum careyi Olney Lycopodium tristachyum Pursh Pinus strobus L. Poa alsodes A. Gray Prunus pensylvanica L. f Panax trifolius L. Pinus virginiana Mill.1 Platanthera clavellata (Michx.) Luer Platanthera lacera (Michx.) G. Don Pyrola elliptica Nutt. Spiranthes ovalis Lindl. var. erostellata Catling Viola blanda Willd. var. palutriformis A. Gray Viola pubescens Aiton 1 Presumed introduced at site 2 Recently documented in Indiana; populations being tracked by IDNR Plant Species of Concern Endangered, threatened and rare plants are tracked by the Indiana Department of Natural Resources (IDNR). Plants on the endangered list have one to five occurrences statewide; those with six to 10 known occurrences are listed as threatened; rare species have 11 to 20 known occurrences. A fourth category of concern is watch list. Watch list species were previously on the ETR list but have since been removed, often because they are represented by greater than 20 occurrences. If a plant is removed from the ETR list, it remains on the watch list perpetually, unless there is reason to move it back to ETR. Watch list species are not actively tracked by the IDNR. (M.A. Homoya, pers. comm., February 2008) Table 4 displays a list of plants observed at Sebert Property that are considered endangered, threatened or rare in Indiana, or that are on the watch list (IDNR 2010). Seven species listed as endangered, three listed as threatened and nine listed as rare were observed on the site. Examples of some of these species can be seen in Figures 10-12. One species previously thought to be extirpated from Indiana, Lonicera canadensis (SAN 152 [MOR]), was observed at the site. Two orchid species that never produced flowers during the study were observed;
Page 11 ï~~2010 THE MICHIGAN BOTANIST 11 2010 THE MICHIGAN BOTANIST 11 FIGURE 10. Indiana watch list species Huperzia lucidula. (Michx.) Trevis. in swamp forest, October 26, 2008. Photograph by Keith Board. FIGURE 11. Indiana watch list species Lycopodium clavatum L. in early successional forest, October 26, 2008. Photograph by Keith Board. FIGURE 12. Indiana rare species Lycopodium tristachyum Pursh in early successional forest, October 26, 2008. Photograph by Keith Board.
Page 12 ï~~12 THE MICHIGAN BOTANIST Vol. 49 TABLE 5. New LaPorte County records (Swink & Wilhelm 1994; The vPlants Project 2009) from Sebert Property. Brachyelytrum aristosum (Michx.) Trel. Carex brunnescens (Pers.) Poir. Carex longii Mack. Cerastium pumilum W. Curtis Corallorhiza odontorhiza (Willd.) Poir. Lonicera canadensis Bartram ex Marsh. Lycopus rubellus Moench. Malus ioensis (Alph. Wood) Britton Pinus virginiana Mill. Populus x canescens (Aiton) Sm. (pro sp.) [alba x tremula ] Ranunculus flabellaris Raf. Salixpetiolaris Sm. Solidago canadensis L. these may be Platanthera psycodes (listed as rare in Indiana; SAN PH-6 [MOR]) and P flava var. herbiola (listed as watch list in Indiana; SAN PH-8 [MOR]). New County Records Table 5 displays a list of 13 plants not previously known from LaPorte County, Indiana (Swink & Wilhelm 1994; The vPlants Project 2009). Of these records, Cerastium pumilum (Board 512 [MOR]), Pinus virginiana (Board 388 [MOR]; SAN 47 [MOR]; SAN 48 [MOR]) and Populus x canescens (Board 422 [MOR]) are introduced; the remaining 10 are native. FIGURE 13. South swamp forest, April 5, 2008. Photograph by Scott Namestnik.
Page 13 ï~~2010 THE MICHIGAN BOTANIST 13 2010 THE MICHIGAN BOTANIST 13 FIGURE 14. South swamp forest, May 31, 2008. Photograph by Scott Namestnik. DISCUSSION Sebert Property consists of a mosaic of plant communities that are collectively considered boreal flatwoods (Figure 2). The most common tree species based on visual observations include Acer rubrum, Betula papyrifera, Fagus grandifolia, Liriodendron tulipifera, Nyssa sylvatica, Quercus alba, Q. bicolor, Q. palustris, Q. rubra, Salix nigra, Sassafras albidum and Ulmus americana. A mix of swamp forest (Figures 13 and 14), mesophytic forest (Figure 15) and upland forest (Figure 16) make up the north and south thirds of the property. These are the communities least disturbed by anthropogenic influences. Tree tip-ups are common throughout the swamp forests (Figure 17), creating functional microhabitats and amphibian breeding pools. Shrub swamp dominated by Cephalanthus occidentalis occupies the northwest corner of the site (Figure 18). Within this buttonbush swamp, a small submerged/emergent wetland that is seasonally to permanently inundated is present in the extreme northwest corner. A somewhat level area that was once a higher sand ridge runs approximately east-west through the middle of the property. As shown on the base map aerial photograph for the LaPorte County Soil Survey, ca. 1970 (Figure 1), this ridge was cleared of vegetation for farming, pasture or potential use as a sand borrow area. These sand flats now exist mostly as disturbed communities including young successional forest (Figure 19) and sand barrens (Figure 20), including a sand blowout area (Figure 21). Conifers in this area that presumably were planted include Pinus sylvestris, P. virginiana, and Thuja occidentalis. Two seasonal ponds that
Page 14 ï~~14 THE MICHIGAN BOTANIST Vol. 49 14 THE MICHIGAN BOTANIST Vol. 49 FIGURE 15. South mesophytic forest, April 5, 2008. Photograph by Scott Namestnik. FIGURE 16. Southeast upland forest, April 5, 2008. Photograph by Scott Namestnik.
Page 15 ï~~2010 THE MICHIGAN BOTANIST 15 2010 THE MICHIGAN BOTANIST 15 FIGURE 17. Tree tip-up in northeast swamp forest, April 5, 2008. Photograph by Scott Namestnik. FIGURE 18. Buttonbush swamp, May 31, 2008. Photograph by Scott Namestnik.
Page 16 ï~~16 THE MICHIGAN BOTANIST Vol. 49 16 THE MICHIGAN BOTANIST Vol. 49 FIGURE 19. Central early successional forest, November 2, 2008. Photograph by Keith Board. can be characterized as acid sand flat openings are present here (Figure 22). Many species in these flats are typically found in artificial excavations where sand has been removed down to the water table (Swink & Wilhelm 1994); it is likely that sand was removed from these areas in the past. Two excavated ditches are present on either side of an abandoned access road at the north end of the site (Figure 23). These ditches lead to the roadside ditch along County Road 925 North (Figure 24); it appears that an attempt was made at some point to drain portions of the property. Comparisons of floristic inventories should be viewed with circumspection, as many variables, including different surveyors, different methods of collecting and identifying species and different levels of survey intensity can all play a part in the number of species observed. With these things in mind, comparing the quality of natural areas in terms of number of species, mean C-values and FQI can generally show where a site ranks in terms of natural area floristic quality. With 434 vascular plants in 40 acres, Sebert Property has a level of species richness greater than that at other sites of equal or greater size for which vascular plant inventories have been recorded in Indiana (Appendix 2) (Rothrock & Homoya 2005; Angstmann et al. 2006; Ruch et al. 2008; Ruch et al. 2008). Of the sites shown in Appendix 2, Barker Woods Nature Preserve is most like Sebert Property in terms of plant communities and species composition. Barker Woods has a higher mean C-value than Sebert Property, but Sebert Property has more than twice the number of species, including natives, resulting in much higher FQI values.
Page 17 ï~~2010 THE MICHIGAN BOTANIST 17 2010 THE MICHIGAN BOTANIST 17 FIGURE 20. West sand barrens, April 5, 2008. Photograph by Scott Namestnik. The numerous plant communities at Sebert Property support a diverse mix of vascular plants, including several ETR species and species with affinities to more boreal or coastal plain regions. It is noteworthy that 83% of the 434 vascular plants documented in the flora of Sebert Property are considered native, and that 13 new county records were discovered. In addition, 31 of the recorded species have C-values of 9 or 10, indicating that portions of the site still harbor pre-settlement natural area quality despite moderate anthropogenic disturbance in the past. Sebert Property is an important tract of land for conservation of at-risk plant species in Indiana. The Lake Michigan-influenced climate, the acidic soils and the location of the site at the northern extent of Indiana allow species that are more common further north to survive here at the southern extent of their geographic ranges. Many of these boreal relicts are considered species of concern in Indiana because they are not known from other parts of the state. The presence of 10 native species not previously known from LaPorte County further illustrates the ecological significance of Sebert Property. The unique assemblage of plant communities, soil types, hydroperiod and disturbance provides appropriate conditions for species otherwise unknown from LaPorte County to survive at this location. This approximately 40-acre tract of land also provides habitat for Lonicera canadensis, a native shrub that was considered extirpated from Indiana prior to this study. Future research at Sebert Property and in northern LaPorte County in general could be directed towards the genera Dichanthelium at drier sites and Betula in forests. A population of an unusual Dichanthelium, determined as D. oligosanthes (SAN 474 [MOR]), was collected in early successional sand barrens at the site. The
Page 18 ï~~18 THE MICHIGAN BOTANIST Vol. 49 18 THE MICHIGAN BOTANIST Vol. 49 FIGURE 21. Central sand barrens, April 5, 2008. Photograph by Scott Namestnik. FIGURE 22. Southeast pond, October 25, 2008. Photograph by Keith Board.
Page 19 ï~~2010 THE MICHIGAN BOTANIST 19 2010 THE MICHIGAN BOTANIST 19 FIGURE 23. Abandoned road, April 5, 2008. Photograph by Scott Namestnik. FIGURE 24. Roadside and roadside ditch, May 31, 2008. Photograph by Scott Namestnik.
Page 20 ï~~20 THE MICHIGAN BOTANIST Vol. 49 plants have long spreading papillose-based upper sheath hairs and purplish culms, and are intermediate between D. oligosanthes ssp. oligosanthes and D. oligosanthes ssp. scribnerianum. Though the plants have longer, narrower leaves characteristic of D. oligosanthes ssp. oligosanthes, they have shorter spikelets characteristic of D. oligosanthes ssp. scribnerianum. Another Dichanthelium of interest, tentatively determined as D. clandestinum x depauperatum (Board 652 [MOR]), was discovered in early successional sand barrens near a sand blowout. Trees of the genus Betula are frequent in northern LaPorte County, and both B. papyrifera and B. populifolia are encountered at Sebert Property. It would be important to determine whether the two are hybridizing, as trees with intermediate characteristics are observed frequently. It is also unclear whether local colonies of B. populifolia are naturally occurring or escaped from cultivation. Perhaps most importantly, many local reports of B. populifolia are misidentifications of B. pendula (European White Birch), which often grows in sandy woods in the dune area, and telling these two species apart can be "quite a trial" (Swink & Wilhelm 1994); this has important implications for management purposes. This study provides a baseline to help future researchers understand the natural and anthropogenic changes occurring at Sebert Property and the surrounding area. Ideally, our study will promote further botanical studies as well as a casual appreciation of the local flora. In addition, it will provide useful data for the owner, LaPorte County Parks Foundation, as management and/or development decisions are considered. To preserve the natural character of the property, it will be increasingly important to monitor the spread of invasive species, which can quickly lead to the degradation of a natural area. This will be especially important along anthropogenically disturbed areas such as the road and trails through the site. Natural succession will also cause changes to the flora of the site. The plant communities present at Sebert Property will require some form of disturbance to be maintained in their present state. In swamp forest areas, windthrown trees naturally open the canopy and reset successional processes. In the acid sand flat openings, historic sand mining removed accumulations of organic material and exposed an ancient seed bank; without disturbance these areas are expected to fill in over time, changing the composition of the acid sand flat openings flora. The successional forest and sand barren communities will also change with time if no management takes place. Future management decisions will determine how the flora of Sebert Property changes in the coming decades. ACKNOWLEDGMENTS The authors wish to thank Tim Morgan, Director of LaPorte County Parks, for permission to conduct this study and for historical information on the property. In addition, valuable assistance with 2005 field work was provided by the following people, and their help is greatly appreciated: Steve Barker, Elizabeth McCloskey, Lindsay Namestnik, Sandy O'Brien, Karen Quinlan, Paul Quinlan and Tony Troche. Finally, the authors wish to express sincere gratitude to the following individuals for detailed review and constructive comments regarding the manuscript: Todd Barkman, Michael Homoya, Bradford Slaughter, Justin Thomas, Dana Thomas and an anonymous reviewer.
Page 21 ï~~2010 THE MICHIGAN BOTANIST 21 LITERATURE CITED Angstmann, J. L., P. E. Rothrock & T. W. Post. (2006). The vascular flora and community structure of Little Calumet Headwaters Nature Preserve, LaPorte County, Indiana. The Michigan Botanist, 45(3): 153-192. Deam, C. C. (1940). Flora of Indiana. Indianapolis, IN: Department of Conservation, Division of Forestry. Fernald, M. L. (1950). Gray's manual of botany: eighth (centennial) edition - illustrated. NY: American Book Company. Flora of North America Editorial Committee, eds. (1993+). Flora of North America North of Mexico. 12+ vols. New York and Oxford. Gleason, H. A. & A. Cronquist. (1991). Manual of vascular plants of northeastern United States and adjacent Canada. New York, NY: New York Botanical Garden. Goff, F. G., G. A. Dawson, & J. J. Rochow. (1982). Site examination for threatened and endangered plant species. Environmental Management, 6: 307-316. Greenberg, J. (2002). A natural history of the Chicago region. Chicago, IL: The University of Chicago Press. Hitchcock, A. S. (1950). Manual of the grasses of the United States. 2nd ed. Revised by Agnes Chase. Reprinted 1971. NY: Dover Publications, Inc. 2 vols. Holmgren, N. H. (1998). Illustrated companion to Gleason and Cronquist's manual. NY: New York Botanical Garden. Homoya, M. A., D. B. Abrell, J. R. Aldrich & T. W. Post. (1985). The natural regions of Indiana. Proceedings of the Indiana Academy of Science, 94: 245-268. Indiana Department of Natural Resources. (2010). Endangered, threatened, rare and extirpated plants of Indiana. Retrieved August 5, 2010, from http://www.in.gov/dnr/naturepreserve/files/np-etrplants 051910.pdf. Jacquart, E., M. Homoya & L. Casebere. (2002). Natural communities of Indiana. Working Draft. Mohlenbrock, R. H. (2002). Vascular flora of Illinois. Carbondale and Edwardsville, IL: Southern Illinois University Press. Rothrock, P. E. (2004). Floristic quality assessment in Indiana: The concept, use, and development of coefficients of conservatism. EPA Wetland Program Development Grant CD975586-01. Rothrock, P. E. & M. A. Homoya. (2005). An evaluation of Indiana's floristic quality assessment. Proceedings of the Indiana Academy of Science, 114(1): 1-8. Ruch, D. G., B. G. Torke, B. R. Hess, K. S. Badger & P. E. Rothrock. (2008). The vascular flora and vegetational communities of the wetland complex on the IMI property in Henry County near Luray, Indiana. Proceedings of the Indiana Academy of Science, 117(2): 142-158. Ruch, D. G., B. G. Torke, K. S. Badger, B. R. Hess, B. N. Christian & P. E. Rothrock. (2008). The vascular flora and vegetational communities of Lick Creek Summit Nature Preserve in Wayne County, Indiana. Proceedings of the Indiana Academy of Science, 117(1): 29-54. Steyermark, J. A. (1963). Flora of Missouri. Ames, IA: The Iowa State University Press. Swink, F. & G. S. Wilhelm. (1994). Plants of the Chicago region. Indianapolis, IN: Indiana Academy of Science. The vPlants Project. (2009). vPlants: A virtual herbarium of the Chicago region. Retrieved December 31, 2009, from www.vplants.org. United States Department of Agriculture, Soil Conservation Service. (1982). Soil survey of LaPorte County, Indiana. USDA, NRCS. (2009). The PLANTS Database. (http://plants.usda.gov, 31 December 2009). National Plant Data Center, Baton Rouge, LA 70874-4490 USA. Voss, E. G. (1972). Michigan flora. Part I: Gymnosperms and monocots. Bloomfield Hills, MI: Cranbrook Institute of Science. Voss, E. G. (1985). Michigan flora. Part II: Dicots (Saururaceae to Cornaceae). Bloomfield Hills, MI: Cranbrook Institute of Science. Voss, E. G. (1996). Michigan flora. Part III: Dicots (Pyrolaceae to Compositae). Bloomfield Hills, MI: Cranbrook Institute of Science. Yatskievych, G. (1999). Steyermark's flora of Missouri: Volume 1. Jefferson City, MO: Missouri Department of Conservation. Yatskievych, G. (2006). Steyermark's flora of Missouri: Volume 2. St. Louis, MO: Missouri Botanical Garden Press.
Page 22 ï~~APPENDICES APPENDIX 1. Vascular flora of Sebert Property. Board = Keith Board; SAN = Scott A. Namestnik; PH = photo documentation. Nomenclature follows The PLANTS Database (USDA, NRCS 2009); where family classifications differ from those of the Angiosperm Phylogeny Group (APG), APG classifications are included in brackets. C-values are from Floristic quality assessment in Indiana: The concept, use, and development of coefficients of conservatism (Rothrock 2004). PTERIDOPHYTES ASPLENIACEAE (Spleenwort Family) Asplenium platyneuron (L.) Britton, Sterns & Poggenb. Ebony Spleenwort BLECHNACEAE (Chainfern Family) Woodwardia virginica (L.) Sm. Virginia Chainfern C=3 Board 596 C=10 Board 701 C=5 SAN 615 DENNSTAEDTIACEAE (Bracken Fern Family) Pteridium aquilinum (L.) Kuhn var. latiusculum (Desv.) Underw. ex A. Heller DRYOPTERIDACEAE (Wood Fern Family) Athyrium filix-femina (L.) Roth Dryopteris intermedia (Muhl. ex Willd.) A. Gray Dryopteris carthusiana (Vill.) H.P. Fuchs Onoclea sensibilis L. Polystichum acrostichoides (Michx.) Schott EQUISETACEAE (Horsetail Family) Equisetum arvense L. LYCOPODIACEAE (Club Moss Family) Huperzia lucidula (Michx.) Trevis. Lycopodiella inundata (L.) Holub Lycopodium clavatum L. Lycopodium digitatum Dill. ex A. Braun Lycopodium obscurum L. Lycopodium tristachyum Pursh Western Brackenfern Common Ladyfern Intermediate Woodfern Spinulose Woodfern Sensitive Fern Christmas Fern C=6 C=10 C=6 C=4 C=5 Board 632 Board 606 Board 586 Board 714 Board 615 n z 0 z H/ Field Horsetail C=1 Board 670 Shining Clubmoss Inundated Clubmoss Running Clubmoss Fan Clubmoss Rare Clubmoss Deeproot Clubmoss Cutleaf Grapefern Rattlesnake Fern C=5 C=7 C=4 C=2 C=6 C=10 SAN 11; SAN 66 SAN PH-s.n. SAN 6 SAN 4 SAN 3 Board 389; Board 542; SAN 7 0 OPHIOGLOSSACEAE (Family) Botrychium dissectum Spreng. Botrychium virginianum (L.) Sw. C=3 SAN PH-3 C=4 Board 507
Page 23 ï~~OSMUNDACEAE (Royal Fern Family) Osmunda cinnamomea L. Osmunda claytoniana L. Osmunda regalis L. var. spectabilis (Willd.) A. Gray THELYPTERIDACEAE (Marsh Fern Family) Thelypteris noveboracensis (L.) Nieuwl. Thelypteris palustris Schott var. pubescens (G. Lawson) Fernald Cinnamon Fern Interrupted Fern Royal Fern New York Fern Eastern Marsh Fern C=9 Board 521 C=8 SAN 286 C=8 Board 543 C=5 Board 642 C=7 Board 685 GYMNOSPERMS CUPRESSACEAE (Cypress Family) Juniperus communis L. Juniperus virginiana L. Thuja occidentalis L. PINACEAE (Pine Family) Pinus strobus L. Pinus sylvestris L. Pinus virginiana Mill. Common Juniper Eastern Redcedar Arborvitae Eastern White Pine Scots Pine Virginia Pine C=10 SAN 128 C=2 SAN 130 ** SAN 108; SAN 127 C=5 SAN 92 * Board 702; SAN 37; SAN 46 ** Board 388; SAN 47; SAN 48 C=5 Board 504; SAN 105 n z 0 z HZ ANGIOSPERMS ACERACEAE (Maple Family) [SAPINDACEAE] Acer rubrum L. ALISMATACEAE (Water Plantain Family) Alisma subcordatum Raf. ANACARDIACEAE (Cashew Family) Rhus copallinum L. var. latifolia Engl. Rhus typhina L. Toxicodendron radicans (L). Kuntze ssp. radicans APIACEAE (Carrot Family) Cicuta maculata L. Daucus carota L. Osmorhiza claytonii (Michx.) C.B. Clarke Red Maple American Water Plantain C=2 Board 677 Winged Sumac Staghorn Sumac Eastern Poison Ivy C= c= c= c= c= Board 659 SAN 302 SAN 453 Spotted Water Hemlock Queen Anne's Lace Clayton's Sweetroot Board 669 Board 684; SAN 41 Board 519
Page 24 ï~~Osmorhiza longistylis (Torr.) DC. Sanicula canadensis L. Sanicula odorata (Raf.) K.M. Pryer & L.R. Phillippe Sium suave Walter Longstyle Sweetroot Canadian Blacksnakeroot Clustered Blacksnakeroot Hemlock Waterparsnip C= C= C= C= Board 520 Board 710; SAN 438 SAN 418 SAN 526 APOCYNACEAE (Dogbane Family) Apocynum cannabinum L. AQUIFOLIACEAE (Holly Family) Ilex verticillata (L.) A. Gray ARACEAE (Arum Family) Arisaema triphyllum (L.) Schott ARALIACEAE (Ginseng Family) Aralia nudicaulis L. Panax trifolius L. Indianhemp C=2 SAN 531 C=8 SAN 434 ASCLEPIADACEAE (Milkweed Family) [APOCYNACEAE] Asclepias incarnata L. Asclepias syriaca L. Asclepias tuberosa L. ASTERACEAE (Aster Family) Achillea millefolium L. Ageratina altissima (L.) King & H. Rob. var. altissima Ambrosia artemisiifolia L. var. elatior (L.) Descourtils Ambrosia trifida L. Antennaria neglecta Greene Antennaria plantaginifolia (L.) Richardson Arctium lappa L. Bidens connata Muhl. ex Willd. Bidens discoidea (Torr. & A. Gray) Britton Bidens frondosa L. Bidens tripartita L. Cichorium intybus L. Conyza canadensis (L.) Cronquist var. canadensis Common Winterberry Jack in the Pulpit Wild Sarsaparilla Dwarf Ginseng Swamp Milkweed Common Milkweed Butterfly Milkweed Common Yarrow White Snakeroot Annual Ragweed Great Ragweed Field Pussytoes Woman's Tobacco Greater Burdock Purplestem Beggarticks Small Beggarticks Devil's Beggartick Threelobe Beggarticks Chicory Canadian Horseweed C=4 Board 487 C=7 SAN 416 C=9 SAN 8; SAN 15 C= c= c= c= c= c= c= c= c= c= c= c= c= c= SAN 546 Board 717 Board 631 n z 0 z H 0q Board 593 Board 722 SAN 570 SAN 579 Board 505 Board 595 SAN 616 SAN 604; SAN 605 SAN 603 SAN 618 SAN 632 Board 719 Board 687
Page 25 ï~~Erechtites hieraciifolia (L.) Raf. ex DC. Erigeron annuus (L.) Pers. Erigeron philadelphicus L. Erigeron strigosus Muhl. ex Willd. Eupatorium perfoliatum L. Eurybia macrophylla (L.) Cass. Euthamia graminifolia (L.) Nutt. Euthamia gymnospermoides Greene Hieracium caespitosum Dumort. Hieracium gronovii L. Hieracium x flagellare Willd. var. flagellare Hypochaeris radicata L. Krigia virginica (L.) Willd. Lactuca biennis (Moench) Fernald Lactuca canadensis L. Leucanthemum vulgare Lam. Prenanthes altissima L. Pseudognaphalium obtusifolium (L.) Hilliard & B.L. Burtt Solidago caesia L. Solidago canadensis L. Solidago juncea Aiton Solidago nemoralis Aiton Solidago rugosa Mill. Sonchus asper (L.) Hill Symphyotrichum dumosum (L.) G.L. Nesom var. dumosum Symphyotrichum lanceolatum (Willd). G.L. Nesom ssp. lanceolatum var. lanceolatum Symphyotrichum lateriflorum (L.) A. Love & D. Love var. lateriflorum Symphyotrichum pilosum (Willd.) G.L. Nesom var. pilosum Taraxacum officinale F.H. Wigg. Tragopogon dubius Scop. Xanthium strumarium L. American Burnweed Eastern Daisy Fleabane Philadelphia Fleabane Prairie Fleabane Common Boneset Bigleaf Aster Flat-top Goldentop Texas Goldentop Meadow Hawkweed Queendevil Hairy Cat's Ear Virginia Dwarfdandelion Tall Blue Lettuce Canada Lettuce Oxeye Daisy Tall Rattlesnakeroot Rabbit-tobacco Wreath Goldenrod Canada Goldenrod Early Goldenrod Gray Goldenrod Wrinkleleaf Goldenrod Spiny Sowthistle Rice Button Aster White Panicle Aster Calico Aster Hairy White Oldfield Aster Common Dandelion Yellow Salsify Rough Cocklebur C= C= C= C= C= C= C= C= c= c= c= * C= c= * C= C= C= * C= SAN 58 SAN 497 Board 558 SAN 437 SAN 571 SAN 597 SAN 566 SAN 600 Board 560 SAN 25 SAN 422 Board 610; SAN 423; SAN 608 Board 495 No Collection Board 691 Board 603 Board 724; SAN 65; SAN 624; SAN 628 SAN 585; SAN 627 SAN 67 Board 725 SAN 541 SAN 69 SAN 70 SAN 511 SAN 54 SAN 50 SAN 64 SAN 68 SAN 161 SAN 506 Board 726 n z 0 z Hq C=2 C=7 0 C=3 C=3 C=6 * C=4 C=3 C=3 C=0 C=0 * C=0 BALSAMINACEAE (Touch-Me-Not Family) Impatiens capensis Meerb. Jewelweed C=2 Board 690
Page 26 ï~~BERBERIDACEAE (Barberry Family) Berberis thunbergii DC. Podophyllum peltatum L. BETULACEAE (Birch Family) Alnus incana (L.) Moench ssp. rugosa (Du Roi) R.T. Clausen Betula papyrifera Marsh. Betula populifolia Marsh. Carpinus caroliniana Walter ssp. virginiana (Marsh.) Furlow Corylus americana Walter Ostrya virginiana (Mill.) K. Koch BRASSICACEAE (Mustard Family) Alliaria petiolata (M. Bieb.) Cavara & Grande Barbarea vulgaris W.T. Aiton Cardamine hirsuta L. Cardamine pensylvanica Muhl. ex Willd. Lepidium campestre (L.) W.T. Aiton Lepidium virginicum L. Rorippa palustris (L.) Besser ssp. fernaldiana (Butters & Abbe) Jonsell CAMPANULACEAE (Bellflower Family) Lobelia cardinalis L. Lobelia inflata L. Japanese Barberry Mayapple Speckled Alder Paper Birch Gray Birch American Hornbeam American Hazelnut Hophornbeam Garlic Mustard Garden Yellowrocket Hairy Bittercress Pennsylvania Bittercress Field Pepperweed Virginia Pepperweed Fernald's Yellowcress * SAN 254 C=3 Board 497 C=9 SAN 111; SAN 159; SAN 288 C=7 Board 633; SAN 109; SAN 126; SAN 157 C=10 SAN 42; SAN 148 C=5 Board 602 C=4 SAN 106 C=5 SAN 432 c= * * c= c= Board 476 Board 477 Board 478 2 Board 488 Board 517 0 Board 713 2 SAN 545 n z 0 z HZ Cardinalflower Indian-tobacco C=4 SAN 499 C=3 SAN 28 CAPRIFOLIACEAE (Honeysuckle Family) Lonicera canadensis Bartram ex Marsh. Sambucus nigra L. ssp. canadensis (L.) R. Bolli Viburnum acerifolium L. Viburnum lentago L. Viburnum opulus L. var. opulus Viburnum prunifolium L. CARYOPHYLLACEAE (Pink Family) Arenaria serpyllifolia L. American Fly Honeysuckle American Black Elderberry Mapleleaf Viburnum Nannyberry European Cranberrybush Blackhaw C= c= c= c= c= SAN 152 No Collection Board 569 SAN 294 SAN 162 SAN 282 0 Thymeleaf Sandwort SAN 297
Page 27 ï~~Cerastium pumilum W. Curtis Cerastium semidecandrum L. Dianthus armeria L. Saponaria officinalis L. Scleranthus annuus L. Silene antirrhina L. Silene latifolia Poir. ssp. alba (Mill.) Greuter & Burdet Stellaria media (L.) Vill. CELASTRACEAE (Staff-Tree Family) Celastrus orbiculatus Thunb. Euonymus obovatus Nutt. CERATOPHYLLACEAE (Hornwort Family) Ceratophyllum demersum L. CHENOPODIACEAE (Family) [AMARANTHACEAE] Chenopodium album L. CLUSIACEAE (Mangosteen Family) [HYPERICACEAE] Hypericum boreale (Britton) E.P. Bicknell Hypericum canadense L. Hypericum majus (A. Gray) Britton Hypericum perforatum L. Hypericum punctatum Lam. Triadenum virginicum (L.) Raf. CORNACEAE (Dogwood Family) Cornus florida L. Cornus obliqua Raf. Nyssa sylvatica Marsh. European Chickweed Fivestamen Chickweed Deptford Pink Bouncingbet German Knotgrass Sleepy Silene Bladder Campion Common Chickweed Oriental Bittersweet Running Strawberry Bush * * * * * * C=0 * Board 512 SAN 298 Board 605 SAN 613 Board 616 SAN 475 SAN 543 Board 482 * SAN 633 C=7 SAN 295 Coon's Tail Lambsquarters C=1 SAN 301 * Board 718; SAN 630 n z 0 z HZ Northern St. Johnswort Lesser Canadian St. Johnswort Large St. Johnswort Common St. Johnswort Spotted St. Johnswort Virginia Marsh St. Johnswort C=8 C=8 C=6 * C=3 C=10 SAN 532 No Collection SAN 39; SAN 559 Board 672 SAN 513 SAN 75; SAN 569; SAN 614 Flowering Dogwood Silky Dogwood Blackgum C= C= C= Board 491 SAN 450 Board 650 CRASSULACEAE (Family) Penthorum sedoides L. Ditch Stonecrop C=2 Board 694 C=3 SAN 580 CUCURBITACEAE (Cucumber Family) Sicyos angulatus L. Oneseed Bur Cucumber
Page 28 ï~~CUSCUTACEAE (Dodder Family) [CONVOLVULACEAE] Cuscuta gronovii Willd. ex Schult. CYPERACEAE (Sedge Family) Carex albicans Willd. ex Spreng. var. emmonsii (Dewey ex Torr.) J. Rettig 00 Scaldweed C=2 Board 682 Emmons' Sedge Carex albolutescens Schwein. Carex arctata Boott ex Hook. Carex blanda Dewey Carex brunnescens (Pers.) Poir. Carex cephalophora Muhl. ex Willd. Carex communis L.H. Bailey Carex comosa Boott Carex crinita Lam. Carex debilis Michx. var. rudgei L.H. Bailey Carex digitalis Willd. Carex folliculata L. Carex gracillima Schwein. Carex granularis Muhl. ex Willd. Carex intumescens Rudge Greenwhite Sedge Drooping Woodland Sedge Eastern Woodland Sedge Brownish Sedge Oval-leaf Sedge Fibrousroot Sedge Longhair Sedge Fringed Sedge White Edge Sedge Slender Woodland Sedge Northern Long Sedge Graceful Sedge Limestone Meadow Sedge Greater Bladder Sedge Spreading Sedge Broad Looseflower Sedge Long's Sedge Hop Sedge Shallow Sedge Muhlenberg's Sedge Pennsylvania Sedge Eastern Star Sedge Rosy Sedge Broom Sedge Weak Stellate Sedge C=5 Board 489; SAN 122; SAN 142; SAN 144; SAN 248; SAN 249 C=9 Board 708 C=10 Board 402; Board 508; SAN 24 C=1 Board 579 C=10 SAN 486 C=3 Board 666 C=8 Board 509; SAN 145 C=6 SAN 424 C=8 Board 580 C=10 SAN 292 C=7 Board 544; Board 581; Board 597 C=10 SAN 32 C=7 Board 536 C=2 Board 555 C=8 Board 598; Board 667; SAN 415; SAN 435 C=7 Board 599 C=7 Board 600; SAN 535 C=5 Board 635; SAN 26 C=4 Board 636; Board 668, Board 680 C=4 SAN 447 C=5 SAN 504 C=5 Board 510 C=4 Board 511 C=5 Board 537 C=4 SAN 29; SAN 568 C=10 Board 539 n z 0 z H 0Z Carex laxiculmis Schwein. var. laxiculmis Carex laxiflora Lam. Carex longii Mack. Carex lupulina Muhl. ex Willd. Carex lurida Wahlenb. Carex muehlenbergii Schkuhr ex Willd. var. muehlenbergii Carex pensylvanica Lam. Carex radiata (Wahlenb.) Small Carex rosea Schkuhr ex Willd. Carex scoparia Schkuhr ex Willd. Carex seorsa Howe
Page 29 ï~~Carex sparganioides Muhl. ex Willd. Carex stipata Muhl. ex Willd. Carex stricta Lam. Carex swanii (Fernald) Mack. Carex tonsa (Fernald) E.P. Bicknell var. rugosperma (Mack.) Crins Carex tonsa (Fernald) E.P. Bicknell var. tonsa Bur-reed Sedge Awlfruit Sedge Upright Sedge Swan's Sedge Parachute Sedge Shaved Sedge Carex tribuloides Wahlenb. Carex virescens Muhl. ex Willd. Carex vulpinoidea Michx. Cyperus lupulinus (Spreng.) Marcks ssp. macilentus (Fernald) Marcks Cyperus strigosus L. Eleocharis engelmannii Steud. Eleocharis obtusa (Willd.) Schult. Fimbristylis autumnalis (L.) Roem. & Schult. Rhynchospora capitellata (Michx.) Vahl Schoenoplectus tabernaemontani (C.C. Gmel.) Palla Scirpus atrovirens Willd. Scirpus cyperinus (L.) Kunth Blunt Broom Sedge Ribbed Sedge Fox Sedge Great Plains Flatsedge Strawcolored Flatsedge Engelmann's Spikerush Blunt Spikerush Slender Fimbry Brownish Beaksedge Softstem Bulrush Green Bulrush Woolgrass C=4 SAN 471 C=2 Board 556 C=5 Board 582 C=4 Board 583 C=9 Board 490; Board 538 C=9 Board 540; SAN 5; SAN 155; SAN 156 C=5 Board 712; SAN 22; SAN 467; SAN 485 C=8 Board 601; Board 637 C=2 SAN 427 C=4 Board 640 C=0 Board 683 C=4 SAN 518; SAN 557 C=1 SAN 599 C=3 SAN 558 C=6 Board 674; SAN 40; SAN 523 C=4 No Collection C=4 SAN 527 C=4 SAN 59 n z 0 z -H ELAEAGNACEAE (Oleaster Family) Elaeagnus umbellata Thunb. Autumn Olive Board 541 ERICACEAE (Heath Family) Gaultheria procumbens L. Gaylussacia baccata (Wangenh.) K. Koch Vaccinium angustifolium Aiton Vaccinium corymbosum L. Vaccinium pallidum Aiton EUPHORBIACEAE (Family) Acalypha rhomboidea Raf. Croton glandulosus L. var. septentrionalis Mull. Arg. Chamaesyce nutans (Lag.) Small Chamaesyce maculata (L.) Small Eastern Teaberry Black Huckleberry Lowbush Blueberry Highbush Blueberry Blue Ridge Blueberry c= c= c= c= c= c= c= c= c= Board 644 SAN 279 SAN 285 Board 501; SAN 141; SAN 146 Board 527 Common Threeseed Mercury Vente Conmigo Eyebane Spotted Sandmat SAN 508 SAN 606 SAN 572 Board 688
Page 30 ï~~FABACEAE (Pea Family) Amphicarpaea bracteata (L.) Fernald var. bracteata Apios americana Medik. Desmodium ciliare (Muhl. ex Willd.) DC. Desmodium glabellum (Michx.) DC. Desmodium nudiflorum (L.) DC. Desmodium paniculatum (L.) DC. Lespedeza hirta (L.) Hornem Medicago lupulina L. Melilotus alba Medikus Robinia pseudoacacia L. Trifolium campestre Schreb. Trifolium pratense L. Vicia villosa Roth American Hogpeanut Groundnut Hairy Small-leaf Ticktrefoil Dillenius' Ticktrefoil Nakedflower Ticktrefoil Panicledleaf Ticktrefoil Hairy Lespedeza Black Medick White Sweetclover Black Locust Field Clover Red Clover Winter Vetch C= C= C= C= C= C= C= * * **C= * *C= * C= C= C= C= C= C= C= SAN 625 SAN 587 SAN 539 SAN 544 SAN 560 SAN 540 SAN 542 Board 693 SAN 446 Board 709 SAN 451 Board 568 Board 619 FAGACEAE (Beech Family) Fagus grandifolia Ehrh. Quercus alba L. Quercus bicolor Willd. Quercus imbricaria Michx. Quercus palustris Miinchh. Quercus rubra L. Quercus velutina Lam. American Beech White Oak Swamp White Oak Shingle Oak Pin Oak Northern Red Oak Black Oak Board 514 Board 657 SAN 31; SAN 507 SAN 564 Board 591;Board 658 SAN 442 SAN 538 n z 0 z HZ GENTIANACEAE (Gentian Family) Bartonia virginica (L.) Britton, Sterns & Poggenb. GERANIACEAE (Geranium Family) Geranium maculatum L. Geranium robertianum L. GROSSULARIACEAE (Gooseberry Family) Ribes cynosbati L. HALORAGACEAE (Water-Milfoil Family) Proserpinaca palustris L. var. crebra Fernald & Grisc. Yellow Screwstem Spotted Geranium Robert Geranium Eastern Prickly Gooseberry C=5 Board 678; SAN 74 C=4 Board 493 C=4 SAN 521 C=4 Board 481 0 Marsh Mermaidweed C=4 SAN 466
Page 31 ï~~HAMAMELIDACEAE (Witch-Hazel Family) Hamamelis virginiana L. IRIDACEAE (Iris Family) Iris virginica L. var. shrevei (Small) E.S. Anderson Sisyrinchium angustifolium Mill. American Witchhazel C=5 Board 516; SAN 53 Shreve's Iris Narrowleaf Blue-eyed Grass C=5 C=3 Board 562 Board 566 JUGLANDACEAE (Walnut Family) Carya cordiformis (Wangenh.) K. Koch Carya ovata (Mill.) K. Koch JUNCACEAE (Rush Family) Juncus acuminatus Michx. Juncus canadensis J. Gay ex Laharpe Juncus effusus L. Juncus marginatus Rostk. Juncus scirpoides Lam. Juncus tenuis Willd. Bitternut Hickory Shagbark Hickory Tapertip Rush Canadian Rush Common Rush Grassleaf Rush Needlepod Rush Poverty Rush C=5 SAN 252 C=4 Board 584 LAMIACEAE (Mint Family) Lamium purpureum L. Lycopus rubellus Moench. Lycopus uniflorus Michx. Monarda punctata L. Prunella vulgaris L. ssp. lanceolata (W. Bartram) Hulten Scutellaria lateriflora L. Stachys tenuifolia Willd. var. hispida (Pursh) Fernald LAURACEAE (Laurel Family) Lindera benzoin (L.) Blume Sassafras albidum (Nutt.) Nees LEMNACEAE (Duckweed Family) [ARACEAE] Lemna minor L. Lemna trisulca L. C= c= c= c= c= c= c= c= c= c= c= c= Board 647 SAN 38; SAN 55 Board 673 No Collection SAN 575 Board 648 Board 479 Board 692; SAN 76 SAN 57; SAN 530 SAN 524 SAN 512 SAN 528 SAN 496 n z 0 z -H Purple Deadnettle Taperleaf Water Horehound Northern Bugleweed Spotted Beebalm Lance Selfheal Blue Skullcap Smooth Hedgenettle Northern Spicebush Sassafras C=5 Board 587; SAN 9; SAN 121 C=1 Board 592 Common Duckweed Star Duckweed C=3 SAN 112 C=6 SAN 93; SAN 113
Page 32 ï~~LILIACEAE (Lily Family) [ALLIACEAE; RUSCACEAE; LILIACEAE] Allium canadense L. Maianthemum canadense Desf. Maianthemum racemosum (L.) Link ssp. racemosum Medeola virginiana L. Polygonatum pubescens (Willd.) Pursh Meadow Garlic Canada Mayflower Feathery False Lily of the Valley Indian Cucumber Hairy Solomon's Seal C= C= C= SAN 429 Board 518 SAN 284 C=8 Board 563 C=4 Board 498 LINACEAE (Family) Linum striatum Walter Ridged Yellow Flax C=5 Board 649; SAN 502; SAN 533 LYTHRACEAE (Loosestrife Family) Lythrum salicaria L. Rotala ramosior (L.) Koehne MAGNOLIACEAE (Magnolia Family) Liriodendron tulipifera L. MALVACEAE (Mallow Family) Abutilon theophrasti Medik. MELASTOMATACEAE (Family) Rhexia virginica L. Purple Loosestrife Lowland Rotala * SAN 629 C=2 SAN 62 Tuliptree Velvetleaf Handsome Harry C=4 Board 588; SAN 436 * No Collection C=5 SAN 583 n z 0 z HZ MONOTROPACEAE (Indian-Pipe Family) [ERICACEAE] Monotropa uniflora L. OLEACEAE (Olive Family) Fraxinus americana L. Fraxinus nigra Marsh. Fraxinus pennsylvanica Marsh. Fraxinus pennsylvanica Marsh. var. subintegerrima (Vahl) Fernald ONAGRACEAE (Evening Primrose Family) Circaea lutetiana L. ssp. canadensis (L.) Asch. & Magnus Indianpipe White Ash Black Ash Red Ash Green Ash C=7 SAN 30 C= c= c= c= c= c= SAN 439 SAN 33; SAN 477 SAN 260 SAN 34 Broadleaf Enchanter's Nightshade Seedbox Board 681 SAN 498 0 Ludwigia alternifolia L.
Page 33 ï~~Ludwigia palustris (L.) Elliot Oenothera biennis L. Oenothera laciniata Hill ORCHIDACEAE (Orchid Family) Corallorhiza odontorhiza (Willd.) Poir. Epipactis helleborine (L.) Crantz Goodyera pubescens (Willd.) R. Br. Platanthera clavellata (Michx.) Luer Platanthera lacera (Michx.) G. Don Liparis liliifolia (L.) Rich. ex Ker Gawl. Spiranthes cernua (L.) Rich. Spiranthes ovalis Lindl. var. erostellata Catling OROBANCHACEAE (Family) Conopholis americana (L.) Wallr. Epifagus virginiana (L.) W. Bartram OXALIDACEAE (Wood Sorrel Family) Oxalis stricta L. PHYTOLACCACEAE (Family) Phytolacca americana L. PLANTAGINACEAE (Plantain Family) Plantago lanceolata L. Plantago rugelii Decne. Marsh Seedbox Common Evening Primrose Cutleaf Evening Primrose Autumn Coralroot Broadleaf Helleborine Downy Rattlesnake Plantain Small Green Wood Orchid Green Fringed Orchid Brown Widelip Orchid Nodding Lady's Tresses October Lady's Tresses American Cancer-root Beechdrops Common Yellow Oxalis C= C= C= SAN 525 SAN 565 SAN 492 C=3 * C=5 C=9 C=4 C=3 C=3 C=3 Board 471 SAN PH-9 SAN PH-2; SAN PH-10 SAN PH-5 SAN PH-7 SAN PH-4 No Collection Board 727 American Pokeweed Narrowleaf Plantain Blackseed Plantain C=8 Board 513 C=8 Board 721 C=0 SAN 296 C=0 SAN 515 * Board 655 C=0 SAN 573 C=3 No Collection n z 0 z HZ PLATANACEAE (Family) Platanus occidentalis L. American Sycamore POACEAE (Grass Family) Agrostis gigantea Roth Agrostis hyemalis (Walter) Britton, Sterns & Poggenb. Agrostis perennans (Walter) Tuck. Andropogon virginicus L. Redtop Winter Bentgrass Upland Bentgrass Broomsedge Bluestem C=2 C=2 C=2 ** Board 630; Board 665; SAN 36 Board 594 SAN 534 SAN 610
Page 34 ï~~Aristida purpurascens Poir. Brachyelytrum aristosum (Michx.) Trel. Brachyelytrum erectum (Schreb. ex Spreng.) P. Beauv. Bromus tectorum L. Calamagrostis canadensis (Michx.) P. Beauv. Cinna arundinacea L. Dactylis glomerata L. Danthonia spicata (L.) P. Beauv. ex Roem. & Schult. Dichanthelium acuminatum (Sw.) Gould & C.A. Clark var. fasciculatum (Torr.) Freckmann Dichanthelium acuminatum (Sw.) Gould & C.A. Clark var. lindheimeri (Nash) Gould & C.A. Clark Dichanthelium clandestinum (L.) Gould Dichanthelium clandestinum x depauperatum Dichanthelium depauperatum (Muhl.) Gould Dichanthelium meridionale (Ashe) Freckmann Dichanthelium oligosanthes (Schult.) Gould Dichanthelium oligosanthes (Schult.) Gould var. scribnerianum (Nash) Gould Dichanthelium sphaerocarpon (Elliot) Gould Digitaria cognata (Schult.) Pilg. Digitaria ischaemum (Schreb.) Schreb. ex Muhl. Digitaria sanguinalis (L.) Scop. Echinochloa crus-galli (L.) P. Beauv. Elymus repens (L.) Gould Eragrostis spectabilis (Pursh) Steud. Festuca ovina L. Festuca subverticillata (Pers.) Alexeev Glyceria septentrionalis Hitchc. Glyceria striata (Lam.) Hitchc. Holcus lanatus L. Leersia oryzoides (L.) Sw. Leersia virginica Willd. Muhlenbergia frondosa (Poir.) Fernald Muhlenbergia mexicana (L.) Trin. Panicum rigidulum Bosc ex Nees Arrowfeather Threeawn Northern Shorthusk Bearded Shorthusk Cheatgrass Bluejoint Sweet Woodreed Orchardgrass Poverty Oatgrass Western Panicgrass Lindheimer Panicgrass Deertongue Starved Panicgrass Matting Rosette Grass Rosette Grass Scribner's Rosette Grass Roundseed Panicgrass Fall Witchgrass Smooth Crabgrass Hairy Crabgrass Barnyardgrass Quackgrass Purple Lovegrass Sheep Fescue Nodding Fescue Floating Mannagrass Fowl Mannagrass Common Velvetgrass Rice Cutgrass Whitegrass Wirestem Muhly Mexican Muhly Redtop Panicgrass C=7 C=10 C=6 * C=5 C=4 C=3 C=3 SAN 72; SAN 607 SAN 487; SAN 522 SAN 536 SAN 491 Board 634; SAN 444 SAN 520; SAN 561 Board 585 Board 641; SAN 440 C=2 Board 653; SAN 23; SAN 609 C=5 Board 654; SAN 470 C=3 Board 651; SAN 431; SAN 596 NA Board 652 C=6 Board 564 C=7 Board 611 C=6 SAN 474 C=6 Board 612 C=5 Board 613 C=2 SAN 490; SAN 567 * SAN 586 * Board 720 * Board 686 * SAN 489 C=3 SAN 71 * Board 608 C=4 Board 607 C=7 Board 671 C=4 Board 609 * Board 646; SAN 27 C=2 SAN 631 C=4 SAN 562 C=3 Board 723 C=4 SAN 590 C=4 SAN 61 n z 0 z H 0q
Page 35 ï~~Paspalum setaceum Michx. var. stramineum (Nash) D. Banks Phalaris arundinacea L. Phleum pratense L. Poa annua L. Poa alsodes A. Gray Poa compressa L. Poa pratensis L. Schenodonorus phoenix (Scop.) Holub Schizachyrium scoparium (Michx.) Nash var. scoparium Setaria faberi Herrm. Setaria pumila (Poir.) Roem. & Schult. ssp. pumila Torreyochloa pallida (Torr.) Church var. pallida Tridens flavus (L.) Hitchc. var. flavus Vulpia octoflora (Walter) Rydb. POLYGALACEAE (Milkwort Family) Polygala polygama Walter POLYGONACEAE (Smartweed Family) Polygonum careyi Olney Polygonum hydropiperoides Michx. Polygonum pensylvanicum L. Polygonum persicaria L. Polygonatum sagittatum L. Polygonum scandens L. Polygonum virginianum L. Rumex acetosella L. Rumex crispus L. Thin Paspalum Reed Canarygrass Timothy Annual Bluegrass Grove Bluegrass Canada Bluegrass Kentucky Bluegrass Tall Fescue Little Bluestem Japanese Bristlegrass Yellow Foxtail Pale False Mannagrass Purpletop Tridens Sixweeks Fescue Racemed Milkwort Carey's Smartweed Swamp Smartweed Pennsylvania Smartweed Spotted Ladysthumb Arrowleaf Tearthumb Climbing False Buckwheat Jumpseed Common Sheep Sorrel Curly Dock C=3 Board 715 * SAN 425 * SAN 441 * Board 523 C=8 Board 522; SAN 256 * Board 614 * Board 524 * Board 559 C=4 SAN 612 * Board 697 * Board 698 C=10 Board 458; Board 645; SAN 443 ** Board 699; SAN 73 C=1 Board 620 C=7 Board 656 n z 0 z HZ C= c= c= * c= c= c= * * SAN 576 Board 696; SAN 35; SAN 63 SAN 588 SAN 577 SAN 589 SAN 578 SAN 519 Board 526 SAN 468 PORTULACACEAE (Purslane Family) Claytonia virginica L. Virginia Springbeauty C=2 Board 472; SAN 16; SAN 125 POTAMOGETONACEAE (Pondweed Family) Potamogeton foliosus Raf. Leafy Pondweed C=4 SAN 510
Page 36 ï~~PRIMULACEAE (Primrose Family) Lysimachia ciliata L. Lysimachia thyrsiflora L. Trientalis borealis Raf. PYROLACEAE (Shin-Leaf Family) [ERICACEAE] Chimaphila maculata (L.) Pursh Pyrola elliptica Nutt. RANUNCULACEAE (Buttercup Family) Actaea pachypoda Elliot Anemone quinquefolia L. Coptis trifolia (L.) Salisb. Ranunculus abortivus L. Ranunculus acris L. Ranunculus flabellaris Raf. Ranunculus recurvatus Poir. ROSACEAE (Rose Family) Agrimonia parviflora Aiton Amelanchier laevis Wiegand Crataegus crus-galli L. Crataegus flabellata (Bosc ex Spach) K. Koch Crataegus mollis Scheele Fragaria virginiana Duchesne Geum canadense Jacq. Malus ioensis (Alph. Wood) Britton Photinia floribunda (Lindl.) K.R. Robertson & Phipps Potentilla simplex Michx. Prunus pensylvanica L. f. Prunus serotina Ehrh. Prunus virginiana L. Rosa multiflora Thunb. Rosa palustris Marsh. Rubus allegheniensis Porter Rubus enslenii Tratt. Fringed Loosestrife Tufted Loosestrife Starflower Striped Prince's Pine Waxflower Shinleaf White Baneberry Wood Anemone Threeleaf Goldthread Littleleaf Buttercup Tall Buttercup Yellow Water Buttercup Blisterwort Harvestlice Allegheny Serviceberry Cockspur Hawthorn Fanleaf Hawthorn Downy Hawthorn Virginia Strawberry White Avens Prairie Crab Apple Purple Chokeberry Common Cinquefoil Pin Cherry Black Cherry Chokecherry Multiflora Rose Swamp Rose Allegheny Blackberry Dewberry C=4 SAN 488 C=7 SAN 426 C=10 Board 500 C=4 Board 639 C=8 SAN 44 C=7 C=7 C=10 C=0 * C=7 C=5 C=4 C=8 C=4 NA C=2 C=2 C=1 C=3 C=8 C=2 C=2 C=1 C=3 * C=5 C=2 C=5 SAN 45; SAN 255 SAN 283 Board 473; SAN 140 Board 480; SAN 12 Board 565 SAN 259 Board 544 SAN 574 SAN 139; SAN 143; SAN 149 SAN 503 SAN 257 SAN 253 Board 492 SAN 430; SAN 469 SAN 563 Board 506 Board 499 SAN 494 Board 590 SAN 258 SAN 147; SAN 428 Board 660 Board 545; SAN 414 Board 525 n z 0 z H 0=
Page 37 ï~~Rubus flagellaris Willd. Rubus hispidus L. Rubus idaeus L. ssp. strigosus (Michx.) Focke Rubus occidentalis L. Rubus pensilvanicus Poir. Sorbus aucuparia L. Spiraea alba Du Roi Spiraea tomentosa L. RUBIACEAE (Madder Family) Cephalanthus occidentalis L. Diodia teres Walter var. teres Galium aparine L. Galium circaezans Michx. var. circaezans Galium circaezans Michx. var. hypomalacum Fernald Galium concinnum Torr. & A. Gray Galium lanceolatum Torr. Galium obtusum Bigelow Galium pilosum Aiton Galium tinctorium (L.) Scop. Galium triflorum Michx. Mitchella repens L. SALICACEAE (Willow Family) Populus deltoides Bartram ex Marsh. Populus grandidentata Michx. Populus tremuloides Michx. Populus x canescens (Aiton) Sm. (pro sp.) [alba x tremula] Salix discolor Muhl. Salix myricoides Muhl. var. myricoides Salix nigra Marsh. Salix petiolaris Sm. SAURURACEAE (Family) Saururus cernuus L. Northern Dewberry Bristly Dewberry Grayleaf Red Raspberry Black Raspberry Pennsylvania Blackberry European Mountain Ash White Meadowsweet Steeplebush Common Buttonbush Poorjoe Stickywilly Licorice Bedstraw Licorice Bedstraw Shining Bedstraw Lanceleaf Wild Licorice Bluntleaf Bedstraw Hairy Bedstraw Stiff Marsh Bedstraw Fragrant Bedstraw Partridgeberry Eastern Cottonwood Bigtooth Aspen Quaking Aspen Gray Poplar Pussy Willow Bayberry Willow Black Willow Meadow Willow C=2 C=6 C=4 C=1 NA C=4 C=4 C=4 C= c= c= c= c= c= c= c= c= c= c= c= SAN 420 Board 661 SAN 617 SAN 413 SAN 417 SAN 280 SAN 516 SAN 517 Board 638 SAN 537 Board 515 Board 689 SAN 476 SAN 500 SAN 433 SAN 419 SAN 493 SAN 445 Board 643 Board 589 n z 0 z Hq C=1 SAN 448 C=4 SAN 129 C=2 SAN 89; SAN 90; SAN 91 * Board 422 C=3 SAN 110; SAN 160; SAN 287 C=9 SAN 124; SAN 158; SAN 250; SAN 601 C=3 SAN 60; SAN 300 C=6 SAN 56; SAN 251 Lizard's Tail C=4 Board 662
Page 38 ï~~SCROPHULARIACEAE (Figwort Family) [PLANTAGINACEAE; LINDERNIACEAE; PHRYMACEAE; SCROPHULARIACEAE] Chelone glabra L. White Turtlehead C=7 SAN Gratiola neglecta Torr. Clammy Hedgehyssop C=4 SAN Gratiola virginiana L. Roundfruit Hedgehyssop C=4 SAN Lindernia dubia (L.) Pennell var. anagallidea (Michx.) Cooperr. Yellowseed False Pimpernel C=2 SAN Lindernia dubia (L.) Pennell var. dubia Yellowseed False Pimpernel C=3 SAN Mimulus ringens L. Allegheny Monkeyflower C=3 SAN Nuttallanthus canadensis (L.) D.A. Sutton Canada Toadflax C=4 Board Verbascum blattaria L. Moth Mullein * SAN Verbascum thapsus L. Common Mullein * SAN Veronica arvensis L. Corn Speedwell * Board Veronica scutellata L. Skullcap Speedwell C=9 Board 00 526 501 43 598 584 502 496 452 505 618 700 SMILACACEAE (Cat-Brier Family) Smilax rotundifolia L. Roundleaf Greenbrier C=4 SAN 281 SOLANACEAE (Nightshade Family) Solanum carolinense L. Solanum dulcamara L. Solanum ptycanthum Dunal SPARGANIACEAE (Bur-Reed Family) [TYPHACEAE] Sparganium americanum Nutt. TILIACEAE (Linden Family) [MALVACEAE] Tilia americana L. ULMACEAE (Elm Family) [CANNABACEAE; ULMACEAE] Celtis occidentalis L. Ulmus americana L. URTICACEAE (Nettle Family) Boehmeria cylindrica (L.) Sw. Pilea pumila (L.) A. Gray Carolina Horsenettle Climbing Nightshade West Indian Nightshade American Bur-reed American Basswood Common Hackberry American Elm Smallspike False Nettle Canadian Clearweed C=0 *C= C=0 Board 716 Board 567 SAN 529 n z 0 z HZ C=10 SAN 495 C=5 Board 711 C=3 No Collection C=3 SAN 107 C=3 Board 679; SAN 52 C=2 Board 695; SAN 51 0 VERBENACEAE (Vervain Family) Verbena hastata L. Swamp Verbena C=3 SAN 509
Page 39 ï~~VIOLACEAE (Violet Family) Viola affinis Leconte Viola blanda Willd. var. palustriformis A. Gray Viola cucullata Aiton Viola labradorica Schrank Viola lanceolata L. Viola macloskeyi Lloyd ssp. pallens (Banks ex Ging) M.S. Baker Viola pubescens Aiton Viola rostrata Pursh Viola sororia Willd. VITACEAE (Grape Family) Parthenocissus quinquefolia (L.) Planch. Vitis aestivalis Michx. Vitis labrusca L. Vitis riparia Michx. Sand Violet Sweet White Violet Marsh Blue Violet Alpine Violet Bog White Violet Smooth White Violet Downy Yellow Violet Longspur Violet Common Blue Violet Virginia Creeper Summer Grape Fox Grape Riverbank Grape C=2 Board 502 C=8 SAN PH-1; SAN 123 C=8 SAN 293 C=7 Board 483 C=9 Board 546 C=10 SAN 10; SAN 154 C=5 Board 474; SAN 13; SAN 151 C=8 Board 475 C=1 Board 484; SAN 14; SAN 150; SAN 153 C=2 SAN 449 C=4 SAN 472; SAN 473 C=1 SAN 421 C=6 SAN 299 Additional taxa not identified to species Platanthera Rich. Platanthera Rich. Typha L. Vernonia Schreb. n z 0 z -H Fringed Orchid Fringed Orchid Cattail Ironweed NA SAN PH-8 NA SAN PH-6 NA No Collection NA No Collection * Non-native ** Native in Indiana (Rothrock 2004), non-native in Chicago Region (Swink & Wilhelm 1994), considered non-native; or obviously introduced at site APPENDIX 2. Published Indiana Floristic Inventories. Total Native Total Native Total Native Site Name, Location Size (ac) Species Species Mean-C Mean-C FQI FQI Barker Woods Nature Preserve, LaPorte County2 29.7 159 146 4.6 5.0 58.4 60.7 Bendix Woods Nature Preserve, St. Joseph County2 Big Oaks National Wildlife Area (3 sample areas)2 Botany Glen, Grant County2 Fall Creek Gorge Nature Preserve, Warren County2 Flint Barrens, Harrison County2 27.2 118 112 51 39 44.5 357 295 148.3 346 288 59.3 289 276 4.4 4.7 48.1 49.3 2.2 2.9 15.8 18.0 3.3 4.0 62.3 68.5 3.1 3.8 58.5 64.1 3.9 4.0 65.6 67.2
Page 40 ï~~Fogwell Forest Nature Preserve, Allen County2 27.2 240 Fox Lake Littoral Zone, Steuben County2 30 Ginn Woods, Delaware County2 150.7 441 Hemlock Bluff Nature Preserve, Jackson County2 39.5 299 IMI Fen, Henry County3 6.2 287 Lick Creek Summit Nature Preserve, Wayne County4 39.0 387 Lime Lake Nature Preserve, Steuben County2 49.4 75 Little Calumet Headwaters Nature Preserve, LaPorte Countyl 108.0 298 Lookout Point Gravel Hill Prairie, Tippecanoe County2 < 2.5 84 Mounds State Park Fen, Madison County2 4.9 97 Mounds State Park Floodplain Forest and Seeps, Madison County2 146 Mounds State Park, Madison County2 259.5 448 Perry County Limestone Glade, Perry County2 < 2.5 47 Perry County Sandstone Glade, Perry County2 < 2.5 32 Plaster Creek Seep Springs, Martin County2 < 2.5 107 Saugany Lake, LaPorte County2 74.1 47 Seasonal Pond and Margin on Taylor University campus, Grant County2 4.9 36 Sebert Property, LaPorte County 39.0 434 Turkey Run State Park Seep Area 1, Parke County2 < 2.5 32 Turkey Run State Park Seep Area 2, Parke County2 < 2.5 43 Twin Swamp Nature Preserve, Posey County2 598.0 138 Upland Prairie Restoration (12th year)2 67 Upland Prairie Restoration (5th year), Grant County2 24.7 65 Versailles State Park Limestone Glade, Ripley County2 < 2.5 83 Wabash Breaks Gravel Hill Prairie, Tippecanoe County2 < 2.5 102 Wea Creek Gravel Hill Prairie, Tippecanoe County2 < 2.5 89 Wening-Sherrit Seep Springs, Dubois County2 < 2.5 93 Wilbur Wright Fish and Wildlife Area2 1028.0 536 Yellow Birch Ravine Nature Preserve, Crawford County2 434.9 420 210 29 364 259 246 304 71 271 74 96 130 380 47 32 107 43 27 362 30 40 125 46 36 80 92 76 90 388 385 3.6 5.2 3.2 3.5 6.0 4.1 4.3 5.9 3.5 3.8 5.2 4.8 5.2 4.4 2.1 3.7 5.1 4.0 3.3 2.5 1.8 4.2 3.9 3.4 5.2 2.8 4.2 4.1 5.4 3.9 4.0 6.3 4.9 5.9 4.0 4.5 5.2 4.8 5.2 4.9 2.7 4.4 5.4 5.1 3.6 3.7 3.2 4.4 4.3 4.0 5.3 3.9 4.6 55.4 59.3 28.7 29.2 67.3 74.1 60.4 64.9 52.0 70.1 39.2 57.7 42.6 80.4 35.7 27.2 53.6 30.5 12.3 76.0 28.8 31.0 38.2 20.8 14.4 38.3 38.9 32.4 49.9 65.8 86.5 53.4 39.2 58.0 45.2 87.3 35.7 27.2 53.6 31.9 14.2 83.3 29.8 32.1 40.2 25.1 19.3 39.0 41.0 35.1 50.7 77.3 90.4 n z 0 z O > 0 Â~ 'Angstmann, J. L., P. E. Rothrock & T. W. Post. (2006). The vascular flora and community structure of Little Calumet Headwaters Nature Preserve, LaPorte County, Indiana. The Michigan Botanist, 45(3): 153-192. 2Rothrock, P. E. & M. A. Homoya. (2005). An evaluation of Indiana's floristic quality assessment. Proceedings of the Indiana Academy of Science, 114(1): 1-8. 3Ruch, D. G., B. G. Torke, B. R. Hess, K. S. Badger & P. E. Rothrock. (2008). The vascular flora and vegetational communities of the wetland complex on the IMI property in Henry County near Luray, Indiana. Proceedings of the Indiana Academy of Science, 117(2): 142-158. 4Ruch, D. G., B. G. Torke, K. S. Badger, B. R. Hess, B. N. Christian & P. E. Rothrock. (2008). The vascular flora and vegetational communities of Lick Creek Summit Nature Preserve in Wayne County, Indiana. Proceedings of the Indiana Academy of Science, 117(1): 29-54.