ï~~144 THE MICHIGAN BOTANIST Vol. 41 The discovery of Eucommia ulmoides as an escape is not necessarily an indication that the species will become invasive. In fact, the species is planted relatively infrequently, and since this is the first report of its escape in the more than 100 years since its introduction in the United States, it seems unlikely that it will become invasive. Hardy rubber-tree does not fit the profile of readily invasive species as described by Reichard & Hamilton (1997). Studies by Call & Dilcher (1997) of the aerodynamics of Eucommia fruits showed that they are dispersed in a downward spiral with an increasing lateral component. The fruits are relatively heavy and not as easily wind-blown as are samaras of other, more invasive woody plants, such as Ulmus species (Anonymous 2002; Sherman-Broyles et al. 1997). If more escapes of this species are to be found, it will likely be in the vicinity of pistillate trees near fence rows and woodlots. LITERATURE CITED Anonymous. 2001. Eucommia bark. http://www.doctorshealthsupply.com/chineseherbs/herbal ingredients/eucommia_bark.htm (accessed 9 October 2002). Anonymous. 2002. Siberian elm (Ulmus pumila). Wisconsin Department of Natural Resources. http://www.dnr.state.wi.us/org/land/er/invasive/factsheets/elm.htm (accessed 6 December 2002). Barker, P.A. 1984. Eucommia ulmoides: a tree for urban areas. Journal of Arboriculture 10: 233-235. Bremer, K., A. Blacklund, B. Sennblad, U. Swenson, K. Andreason, M. Hjerston, J. Lundberg, M. Blacklund, & B. Bremer. 2001. A phylogenetic analysis of 100+ genera and 50+ families of euasterids based on morphological and molecular data with notes on possible higher morphological synapomorphies. Plant Systematics and Evolution 229: 137-169. Call, V.B. & D.L. Dilcher. 1997. The fossil record of Eucommia (Eucommiaceae) in North America. American Journal of Botany 84: 798-814. Coombes, A.J. 1992. Eyewitness Handbooks of Trees. Dorling Kindersley, New York, NY. 320 pp. Cronquist, A. 1988. The Evolution and Classification of Flowering Plants (ed. 2). New York Botanical Garden, Bronx, NY. 555 pp. Dirr, M.A. 1990. Manual of Woody Landscape Plants: Their Identification, Ornamental Characteristics, Culture, Propagation and Uses. Stipes, Champaign, IL. 1007 pp. Dirr, M.A. 1997. Dirr's Hardy Trees and Shrubs. Timber Press, Portland, OR. 493 pp. Flint, H.L. 1983. Landscape Plants for Eastern North America. Wiley & Sons, New York, NY. 677 pp. Forrest, T. 1995. Two thousand years of eating bark: Magnolia officinalis var. biloba and Eucommia ulmoides in traditional Chinese medicine. Arnoldia 55: 13-18. Gilman, E.F. 1997. Trees for Urban and Suburban Landscapes. Delmar, Albany, NY. 662 pp. Gilman, E.F. & D.G. Watson. 1993. Eucommia ulmoides-Hardy Rubber Tree. Environmental Horticulture Department, Florida Cooperative Extension Service, Institute of Food and Agricultural Sciences, University of Florida. Fact Sheet ST-240. 3 pp. Harms, H. 1930. Eucommiaceae. In: Engler, A. & K. Prantl, Die Naturlichen Pflanzenfamilien, Verlag W. Engelmann, Leipzig. 18a: 348-351. Hutchinson, J. 1967. The Genera of Flowering Plants (Angiospermae). Clarendon Press, Oxford. 2 vols. Jeon, H.-J., J.-Y. Roh, D.-H. Kim, W.-K. Lee, & Y.-J. Ahn. 1998. Growth-inhibiting effects of various traditional drinks of plant origin on human intestinal bacteria. Hanguk Nongwhahak Hoechi 41(8): 605-607. Kubitzki, K. 1993. The Families and Genera of Vascular Plants. Vol. 2. Flowering Plants-Dicotyledons. Magnoliid, Hamamelid and Caryophyllid Families. Springer-Verlag, Berlin. 653 pp. Li, Y., T. Sato, K. Metori, K. Koike, Q.M. Che, & S. Takahashi. 1998. The promoting effects of geniposidic acid and aucubin in Eucommia ulmoides Oliver leaves on collagen synthesis. Biological and Pharmaceutical Bulletin 21(12): 1306-1310. Mabberley, D.J. 1997. The Plant-Book (ed. 2). Cambridge University Press, Cambridge, UK. 858 pp.
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