The North American species of Psathyrella.

1972]

PSATHYRELLA

93

5. Not as above. 6
6. Pileus viscid, whitish with a lavender tinge; pileus "radially floccose-striate".
57. P. elongatipes.
6. Pileus not viscid. 7
7. Pileus pallid (whitish) at first. 58. P. huronensis.
7. Pileus yellowish to gray-brown to vinaceous-brown. 8
8. Spores truncate at apex; growing cespitose on hardwood stumps and logs.
59. P. waltersii.
8. Spore apex not truncate. 9
9. Grayish tones evident on young pileus; solitary to gregarious under aspen in the
Rocky Mountains. 60. P. uliginicola.
9. Pileus pale yellow when young; on grassy ground, Tennessee. 61. P. luteopallida.
52. Psathyrella longipes (Peck) A. H. Smith, Contr. Univ. Mich. Herb. 5: 49.
1941.
Hypholoma longipes Peck, Bull. Torrey Club 22: 204. 1895.
Hypholoma campanulatum Peck, Bull. Torrey Club 36: 336. 1909.
Drosophila campanulata (Pk.) Murrill, Mycologia 4: 304. 1912.
Illust. Text Figs. 134-136.
Pileus 2.5-3.7 cm broad, campanulate, even or obscurely striate on the margin,
fibrillose becoming glabrous, hygrophanous, yellowish-brown when moist, brown
or isabelline-brown when dry, the margin appendiculate with the very white
floccose, fugaceous veil. Context extremely thin and fragile.
Lamellae narrow, close, adnate, white or whitish becoming nearly black, often
whitish on the edge.
Stipe slender, 5-10(-12.5) cm long, 2-5 mm thick, hollow, striate at the top,
white with a white mycelioid tomentum at the base.
Spores 10-13 x 7-8x 8-10 /, smooth, apical pore distinct, shape in face view
ovate to elliptic and (as revived in KOH) showing a tendency to angularity, in
profile subelliptic to obscurely inequilateral, color in KOH at first mummy brown
slowly becoming dark chocolate-color, in Melzer's tawny-red, wall about 0.8-1,u
thick (in KOH) in old spores.
Basidia 4-spored, 16-22 X9-10 A, clavate, hyaline in KOH. Pleurocystidia
none. Cheilocystidia 32-48x9-15 /, fusoid-ventricose with obtuse apex varying
to narrowly ventricose to clavate, thin-walled, smooth, hyaline, content not
distinctive. Brachybasidioles present (use mature pilei). Caulocystidia scattered
to numerous, clavate, to fusoid-ventricose or utriform, variable in size, walls thin,
smooth and hyaline, cell content not distinctive.
Pileus with a cuticle of vesiculose cells several deep, the walls thin, smooth
and hyaline, the content of the cells not distinctive. Hyphae of the trama hyaline
in KOH, thin-walled, walls smooth. No distinctive color reactions in any tissue
as revived in Melzer's. Clamp connections present.
Type locality. Pasadena, Calif.
Habit and habitat. Among fallen leaves in very wet weather, late fall or
winter.
Distribution. California, type studied, Michigan, Oregon, Washington.
Observations. Smith (1941) misidentified collections from the Olympic Mountains of Washington as this species. For an account of this material see P.
elwhaensis. The latter does not have flattened spores. The spore size and shape
for H. campanulatum are the same as for P. longipes. The distinguishing features
of the latter as based on a study of the type are as follows: the spores are
compressed and somewhat angular in face view, pleurocystidia are lacking, and