The North American species of Psathyrella.

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MEMOIRS OF THE NEW YORK BOTANICAL GARDEN

[VOL. 24

appendiculatus; sapor subnauseosus; lamellae pallidae demum purpureo-brunneae,
adnatae secedentes latae confertae; stipes 6-7 cm longus, 5-7 mm crassus; deorsum
subsquamulosus et demum sordide brunneus; sporae 8-11 x 4-4.5 /A; fibulae adsunt.
Typus. Smith 55708 (MICH); legit prope Grants Pass, Oregon.
Illust. P1. 27, fig. a; Text Figs. 127, 128.
Pileus 4-6 cm broad, convex becoming plane, surface dark honey-color to near
"buckthorn brown," fading on disc first to a dingy warm buff, when young coated
with "pinkish buff" fibrils as remnants of an outer veil, or squamulose over all,
margin appendiculate with remains of a whitish veil. Context thin but firm, taste
subnauseous, odor none.
Lamellae pallid then grayish and finally purple brownish, adnate, seceding,
broad, close, edges even.
Stipe 6-7 cm long, 5-7 mm thick, equal, pallid, surface fibrillose to squamulose
over lower two-thirds, base brownish where handled, apex striate and pruinosescurfy.
Spores 8-11x 4-4.5 au, smooth, apical pore broad, shape in face view elliptic
to subovate, in profile obscurely bean-shaped to obscurely inequilateral, color in
KOH fuscous (a weak ochraceous tone present), only slowly changing to chocolate
color in KOH, in Melzer's tawny-red, wall 0.3 / thick.
Basidia 4-spored, 24-30x7-11 /u, clavate, hyaline in KOH. Pleurocystidia
very rare and found only near the gill edge, similar to the cheilocystidia. Cheilocystidia 28-34 x 7-13 [, clavate to utriform, thin-walled, hyaline in KOH, content
"empty" in either KOH or Melzer's. Caulocystidia near stipe apex, mostly clavate
but varying to utriform, sparse to absent lower down.
Pileus cuticle of inflated cells 2-4 deep, the walls thin and hyaline to yellowish
hyaline in KOH, content not distinctive, subcutis and tramal hyphae not appreciably colored in KOH or very soon fading if colored at first; hyphae of veil yellow
in KOH. Clamps present. No distinctive reactions on any tissue observed in
mounts revived in Melzer's.
Type locality. Grants Pass, Oregon, near Rogue River.
Habit and habitat. Scattered on clay soil in an old road through mixed woods,
November.
Distribution. Known only from type locality.
Observations. This species is obviously in the stirps Candolleana in spite of
the occurrence of a few cystidia near the gill edge. It differs from P. candolleana
in the relatively strong pigmentation of the pileus ("buckthorn brown" or darker),
the pinkish buff veil remnants which under the microscope in KOH are yellow
instead of pallid, the stipe which discolors over the basal area from handling, and
the occurrence of veil remnants in patches and squamules over the lower half of
the stipe.
50. Psathyrella hymenocephala (Peck) A. H. Smith, Contr. Univ. Mich. Herb.
5: 47. 1941.
Agaricus hymenocephalus Peck, Ann. Rep. N. Y. State Mus. 31: 34. 1879.
Hypholoma hymenocephalum (Pk.) Saccardo, Sylloge Fung. 5: 1042. 1887.
Hypholoma cinereum Parker, Mycologia 25: 175. 1933.
Atylospora striatula Murrill, Mycologia 14: 267. 1922.
Psathyra striatula (Murrill) Murrill, Mycologia 14: 278. 1922.
Illust. P1. 28; Text Figs. 129, 130.
Pileus (1-) 2-4 (-8) cm broad, obtusely conic to convex when young, the margin
straight, in age becoming broadly convex to plane or somewhat umbonate, surface