The North American species of Psathyrella.

1972]

PSATHYRELLA

297

features large spores, utriform pleurocystidia, a rudimentary grayish fibrillose
veil and no brown-walled setae on the pileus. As far as is known there is no
pseudorhiza.
Material examined. California: Lucke, Dec. 1933. New York: Peck's type.
264. Psathyrella pseudolarga A. H. Smith, sp. nov.
Pileus 4-9 cm latus, convexus vel planus, rugulosa, glaber; lamellae triste
vinaceo-brunneae, latae, confertae; stipes 8-14 cm longus, 6-12 mm crassus,
albidus, sparse fibrillosus; velum nullum; sporae (8-)9-12x (4.5-)5-6.5(-7) pt;
pleurocystidia 42-60x 10-18,u, utriformia; fibulae adsunt. Typus. Smith 55955
(MICH); legit prope Crescent City, California.
Illust. P1. 76; Text Figs. 559-561.
Pileus 4-9 cm broad, obtuse to convex, expanding to plane, surface glabrous,
moist, hygrophanous, radially wrinkled, "warm sepia" (dark cinnamon), fading
to pinkish buff. Context very thin and fragile, odor and taste not distinctive.
Lamellae when young grayish brown, in age dark vinaceous-brown ("Natal
brown"), broad, adnate, seceding, close, edges even and not beaded with drops.
Stipe 8-14 cm long, 6-12 mm thick, equal, fragile, hollow, whitish over all
inside and out, surface appressed fibrillose below and silky above; no veil evident
even on the button stages.
Spores (8-)9-12x (4.5-)5-6.5(-7) 1u, variable in size, smooth, apical pore
distinct, shape in face view ovate to elliptic, in profile somewhat bean-shaped to
obscurely inequilateral, dingy cocoa-color when revived in KOH and becoming
paler on standing, in Melzer's reddish tawny.
Basidia 4-spored, 8-10 /u broad, 20-25 /u long. Pleurocystidia 42-60 x 10-18 /,
broadly utriform, apex subcapitate in many, wall 0.2-0.5,u thick and hyaline to
yellowish in KOH, smooth, content not distinctive in KOH or in Melzer's. Cheilocystidia vesiculose to clavate or subutriform (neck poorly developed), edge
ochraceous from collapsed cells (color in the wall but dilute to the degree that
isolated cells appear hyaline). Caulocystidia rare to scattered, clavate to vesiculose or varying to utriform, very delicate and reviving poorly (apparently there
is a mixture of pleuro- and cheilocystidial types).
Pileus with a cuticle of inflated cells 2-3 deep, the long axis of the cells
perpendicular to the pileus surface, walls (double-wall) about 1, thick, hyaline
in KOH; subcutis and context hyphae weakly brown in KOH and becoming paler,
walls thin and smooth, hyaline lens-shaped wall thickening present erratically in
the tissues of the basidiocarp. Clamp connections present. No distinctive reaction
observed on any tissue as mounted in Melzer's.
Type locality. Crescent City, California.
Habit and habitat. On a log of Madroia, November.
Distribution. Known only from type locality.
Observations. This species is closely related to P. larga but differs in the larger
spores, in the absence of a veil and in that the color of the spores become paler in
KOH rather than darker on standing (for a short time).
265. Psathyrella galerinoides A. H. Smith, sp. nov.
Pileus 2-5 mm latus campanulatus, glaber, griseo-brunneus, mollis; lamellae
latae, distantes pallidae demum violaceo-brunneae; stipes 1-2 longus, circa 0.5 mm
crassus, mollis, pruinosus, aquose cinereus; sporae 8-11x5-6 /; pleurocystidia