The North American species of Psathyrella.

1972]

PSATHYRELLA

295

Lamellae broad, adnate, subdistant, becoming horizontal, dull rusty brown
and drying this color, edges even.
Stipe 1-2 cm long, about 1 mm thick, equal, delicate, pallid, thinly coated
at first with pallid fibrils from the remains of a thin veil, not discolored appreciably in age.
Spores 11-14(-15) x 5.5-7 t, smooth, apical pore indistinct and apex rounded,
shape in face view mostly ovate, in profile somewhat inequilateral, color in
KOH dull cocoa-color to fuscous (with an obscure ochraceous tint), slowly
becoming dark chocolate-color, in Melzer's reddish tawny, wall about 0.3 p.
thick.
Basidia 4-spored, clavate, 18-26 x 8-10 /, hyaline in KOH or pedicel weakly
cinnamon colored. Pleurocystidia scattered, 37-48 (-56) x 10-16 /u, fusoid-ventricose, apex obtuse to rounded, wall thin, smooth and hyaline, or a few with a
granular incrustation over the apex, content not distinctive in either KOH or
Melzer's. Cheilocystidia 30-40 x 10-15,, broadly ventricose, neck short and
apex obtuse to rounded, wall and content as in pleurocystidia. Caulocystidia
clavate, scattered, reviving poorly, walls weakly ochraceous in KOH. Hyphae of
cortex weakly cinnamon in mass and with cinnamon debris in places, much
granular material present in KOH mounts and some adhering to the hyphae.
Cuticle of pileus a layer of vesiculose cells 1-2 deep, the walls weakly
colored cinnamon or more ochraceous in KOH, thin and smooth or nearly so.
Hyphae of pileus trama cinnamon in KOH and with some incrusting material on
the narrower hyphae. Clamps present. No distinctive reactions on any tissue in
Melzer's.
Type locality. Mt. Rainier National Park, Washington.
Habit and habitat. Scattered on conifer needles (a mixed stand of hemlock,
Abies and Pseudotsuga).
Distribution. Known only from type locality.
Observations. Psathyrella pseudofrustulenta is distinguished by the grayish
veil fibrils, rusty brown color of pileus reminding one of P. frustulenta, broad
subdistant lamellae, large spores somewhat inequilateral in profile view, and
rather "average" pleurocystidia for the genus. The lack of a distinct germ pore
in a spore of this size is also unusual in this genus.
262. Psathyrella pseudofoenisecii A. H. Smith, sp. nov.
Pileus 1-3 cm latus, conicus, glaber, purpureo-brunneus demum pallidus dein
ad marginem incarnatus; lamellae latae, subdistantes, brunneolae, demum fuscae;
stipes 4-7 cm longus, 2.5-3.5 cm crassus; pallidus ad basem albo-myceliosus,
tactu deorsum vinaceo-brunneus; sporae 12-16 x 6-7.5 p.; pleurocystidia 38-57 x
12-18 p/, fusoide ventricosa, obtusa vel rotundata; fibulae adsunt. Typus. Nimke
99 (MICH); legit Ann Arbor, Michigan.
Pileus 1-3 cm broad, obtusely conic becoming broadly conic, margin straight
when young, surface glabrous, dingy purplish brown moist, pallid (whitish)
when faded and later becoming flushed pink near the margin; no veil present.
Context very fragile, thin, pallid when faded, odor and taste not distinctive,
FeSO, no reaction.
Lamellae ascending adnate, broad, subdistant, "wood brown" when mature,
merely pallid brownish young, edges even and pallid.
Stipe 4-7 cm long, 2.5-3.5 mm thick, equal, hollow, fragile, pallid overall,