The North American species of Psathyrella.

152 MEMOIRS OF THE NEW YORK BOTANICAL GARDEN [VOL. 24
to cystidioid end-cells. Hyphae of the stipe cortex with hyaline to weakly
ochraceous walls in KOH, hyphae of the veil on the stipe with ochraceous-tawny
coagulated content or wall-color and the walls also minutely roughened at times.
Gill trama parallel or nearly so, tinged cinnamon revived in KOH but fading,
hyaline in mounts of fresh material in HO2 or KOH. Pileus with a cuticle
consisting of a layer of elongated (perpendicularly to pileus surface) or clavate
to vesiculose cells about 2 cells deep, walls ochraceous-cinnamon as revived in
KOH (at first dark cinnamon-brown), the pedicels of the clavate cells tawny and
with thickened walls, in water mounts of fresh material the surface-cells yellowish
or in KOH cinnamon. Hyphae of the context cinnamon in KOH as revived but
fading on standing, nearly hyaline in H20 fresh, walls smooth to minutely roughened. Clamp connections present. All tissues as revived in Melzer's pale greenish
yellow.
Type locality. Eagle Peak, Mt. Rainier National Park, Washington.
Habit and habitat. On soil forming face of a bank along a trail, conifer
roots were close by.
Distribution. Washington.
Observations. The colored walls of the cuticular hyphae make this one of
an outstanding group occurring on conifer wood.
Material examined. Washington: Smith 30166, 30646, 31105 (Type).
107. Psathyrella sequoiae Thiers & A. H. Smith, sp. nov.
Pileus 2.5-4 cm latus, demum late convexus, sparse fibrillosus, glabrescens, ad
marginem appendiculatus, cinnamomeo-brunneus; lamellae demum subdistantes,
latae, griseae demum subspadiceae; stipes 4-6 cm longus, 3-8 mm crassus; albidus,
deorsum tactu brunneus, glabrescens; sporae 6-7.5 x 3-3.5 x 3.6-4 (-4.5) p; pleurocystidia 34-48x 10-15x7-11 u, lecythiformia; fibulae adsunt. Typus. Thiers
18282 (MICH); Del Norte County, California.
Pileus 2.5-4 cm broad, subglobose to convex when young, typically broadly
convex at maturity, surface at first thinly fibrillose from remnants of the outer
veil, more densely white-fibrillose along the appendiculate margin, glabrous in
age or remnants of the veil remaining along the margin, color dark rusty brown
(near "cinnamon-brown"), when dried dark reddish brown on mature pilei.
Context fragile, concolorous with surface, odor and taste not distinctive.
Lamellae adnate to adnexed, close to subdistant, thin, fragile, several tiers of
lamellulae; drab when young, to buffy brown when older but cocoa-brown as
dried, edges entire, paler in color.
Stipe 4-6 cm long, 3-8 mm thick, equal, white, staining brown around base
where handled, dry, lower portion white-fibrillose from veil remnants variously
arranged, with a slight annular fibrillose zone at times where the veil breaks,
silky near the apex.
Spores 6-7.5x3-3.5x 3.6-4(-4.5) j/, smooth, apex truncate in many from a
distinct hyaline pore, shape in face view broadly oval to elliptic, in profile
somewhat bean-shaped to obscurely inequilateral (very slightly compressed),
color in KOH cocoa-color, slowly graying slightly, in Melzer's pale tawny, wall
about 0.2,u thick.
Basidia 4-spored, narrowly clavate, 15-19 x 6-7 /, hyaline. Pleurocystidia
scattered 34-48 x 10-15 x 7-11, nine-pin-shaped, walls thin, smooth and hyaline,
cell content not distinctive. Cheilocystidia similar to pleurocystidia or with
apex merely obtuse and many more nearly clavate-mucronate than nine-pinshaped, smooth, hyaline and thin-walled.