The North American species of Psathyrella.

1972]

PSATHYRELLA

97

subdistant, one tier of lamellulae, pallid brownish young, dark reddish brown in
age, edges even.
Stipe 2-3 cm long, 4-6 mm thick, equal or base bulbous from adhering sand,
not fragile, stuffed but soon hollow, cortex dull brownish, surface whitish to pallid
and at first fibrillose with the remains of the veil, apex often striate.
Spores 9-11 x 5-6.5 /u, smooth, apical pore broad and shallow but apex rounded,
shape in face view elliptic to broadly elliptic, in profile subelliptic, color in KOH
reddish chocolate-color darkening to dark chocolate-color, in Melzer's tawny-red,
wall about 0.3-0.4 K thick.
Basidia 28-34 x 8-12 /, 4-spored, hyaline in KOH, clavate. Pleurocystidia
absent (a few near the edge and resembling cheilocystidia). Cheilocystidia
28-42 x 14-18 /, vesiculose, clavate or broadly fusoid-ventricose with rounded
apex, walls thin, smooth and hyaline, cell content not distinctive, a few with
slightly thickened (0.4 /) yellowish walls in KOH. Caulocystidia vesiculose to
clavate or broadly ventricose, some up to 30 /u broad, abundant, walls thin, smooth
and hyaline. Hyphae of the cortex mostly with hyaline wall thickenings of a
local type sufficiently numerous to cause the hyphae to appear to have roughened
walls from the pattern in which the light is refracted.
Gill trama of regularly arranged hyphae (slightly interwoven), very pale
brownish in water mounts of fresh material, cocoa-color to yellow-brown as
revived in KOH, the walls with incrustations. Pileus cuticle mostly of pearshaped to clavate cells vertically arranged into more or less of a palisade, the
walls hyaline or only the base slightly colored, cell content not distinctive.
Hyphae of the trama dark cocoa-colored when first revived in KOH, fading to
a dingy yellow-brown, the hyphae coarsely incrusted with amorphous colored
material (in KOH), in Melzer's the incrusting material colorless. Clamp connections present.
Type locality. South Lyon, Michigan.
Habit and habitat. Gregarious on waste soil, often in sand.
Distribution. Kansas and Michigan.
Observations. The medium-large spores, numerous and greatly enlarged
caulocystidia, habitat, and heavily incrusted (for the genus) tramal hyphae
make this an easily recognized species.
Material examined. Kansas: Brooks 1604. Michigan: Smith 18929 (Type),
18930.
57. Psathyrella elongatipes (Parker) A. H. Smith, comb. nov.
Hypholoma elongatipes Parker, Mycologia 25: 196. 1933.
Hypholoma longipes Dearness & Bisby, Fungi of Manitoba, 113. 1929 (non H.
longipes Peck).
Illust. Text Figs. 147, 148.
Pileus 6-14 cm broad, globose then hemispheric and finally plano-convex or
at times the disc broadly umbonate and the margin slightly turned up, radiately
floccose-striate especially towards the margin, viscid, whitish with a lavender
tinge, becoming buff, drying buff to buckthorn brown. Context whitish, thin to
the umbo, taste and odor mild.
Lamellae broad, chocolate brown at maturity but whitish at first and soon
purplish and mottled, adnate or arcuate and adnate-decurrent by ridges on the
stipe, edge white-fimbriate, not distilling droplets.
Stipe 6-18 cm long, 1-3 cm thick, shining, concolorous or paler than the
pileus, sometimes brownish purple at the base, even, innately fibrillose, striate,