The veiled species of Hebeloma in the western United States / Alexander H. Smith, Vera Stucky Evenson, and Duane H. Mitchel.

The Subgenus Hebeloma

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23. Hebeloma sublamellatum sp. nov.
Pileus 2.4-4 cm latus, late convexus, glaber, ad centrum obscure
rubrobrunneus, ad marginem griseobrunneus, saepe virgatus, demum
brunneomaculatus. Velum sparsim, fibrillosum. Odor et gustus raphaninus. Lamellae pallidae demum obscure rubrobrunneae, confertae, latae, adnatae. Stipes 3-6 cm longus, 4-9 mm crassus, ad basin
brunneus. Sporae 8-10 (11) x 5-6 pLm, phaseoliformes vel ovoideae
vel oblongae, subleves. Cheilocystidia 52-68 x 7-9 x 4-6 JIm, anguste fusoid-ventricosa.
Specimen typicum in Herb. Univ. Mich. conservatum est, Smith
87044; legit prope Independence Pass, Pitkin County, Colorado, 4 Aug
1976, Mrs. Robert Scates.
Pileus 2.4-4 cm broad, broadly convex, the margin inrolled,
surface "Verona Brown" on disc and grayish brown over the marginal area which is often virgate, tending to stain bister or darker over
disc in age, margin decorated with veil remnants for only a short
time. Veil + pinkish buff. Context watery brown, fading to white;
odor and taste raphanoid, FeSO4 greenish on the stipe, KOH no
reaction.
Lamellae moderately broad, close, adnexed, edges even, surface
pallid then pinkish brown ("Verona Brown"), not staining.
Stipe 3-6 cm long, 4-9 mm thick, equal, solid, pallid within but
brownish in the base; surface with patches of the buff-colored veil
variously disposed, cortex slowly darkening from the base upward.
Spores 8-10 (11) x 5-6 pIm, in profile obscurely bean-shaped to
oblong, in face view elliptic to ovate, appearing smooth under a highdry lens, in KOH brownish, not dextrinoid.
Hymenium.-Basidia 4-spored, 10-12 jLm broad near apex. Pleurocystidia none. Cheilocystidia 52-68 x 7-9 x 4-6 (xm, ventricose
near base and with an elongated neck.
Lamellar and pilear tissues.-Lamellar trama typical for the genus. Cuticle of pileus an ixocutis, the hyphae 2-3 jLm diam and yellowish in KOH, clamp connections present. Hypodermium a thick layer of
rusty brown cells as revived in KOH.
Habit, habitat, and distribution.-Scattered under conifers, Independence Pass, Pitkin County, Colorado, August 4, 1976; Mrs. Robert
Scates (type, MICH, Smith 87044).
Observations.-This species is distinct from H. luteobrunneum in
having an ixocutis over the pileus, in having a raphanoid odor and
taste, and in having a pileus which becomes ~ spotted darker in age. It
may eventually be found desirable to regard it as a variety of H.
mesophaeum.