North American species of Lactarius / L. R. Hesler, Alexander H. Smith.

Subgenus Tristes

391

icum in Herbarium University of Tennessee conservatum, Bigelow
12432 (TENN 35341), South Conway, New Hampshire, 4 Sep 1963.
Pileus 3-7 cm broad, convex to plane, "hair-brown" to "benzobrown," appearing frosted, becoming near "light mouse gray" when
dried, viscid, and somewhat shiny, azonate, matted-fibrillose (under a
lens). Context mild. Latex white, unchanging, not staining the lamellae,
taste mild.
Lamellae subdecurrent, dull "cartridge-buff," close, broad, lamellulae rather numerous.
Stipe 3.5-6.5 cm long, 12-15 mm thick, "light drab," appearing
frosted, finally matted-fibrillose (under a lens), viscid (?), tapering
downward, hollow.
Spore deposit "cartridge-buff" (pale yellow). Spores (from sections) 7-8 X 6-7,p, broadly ellipsoid, plage very small or absent; ornamentation of unbranched to sparsely branched ridges forming a broken
reticulum at times (entire wall somewhat amyloid); prominences 0.5-1
(1.5) / high.
Basidia about 45 X 9,, 4-spored. Pleurocystidia: macrocystidia
60-75 X 6-9/, fusoid to fusoid-ventricose and acute, some aciculate,
some with one or more subapical constrictions; pleurocystidia not observed. Cheilocystidia 37-60 X 4.5-7.5 ui, subclavate to subfusoid,
rounded becoming acute. Gill trama lacking rosettes, lactifers inconspicuous. Pileus trama heteromerous, lactifers inconspicuous, dextrinoid debris present in upper trama and subcutis. Pileus cuticle poorly
delimited as a cutis of loosely tangled hyphae in a slime matrix (an ixocutis), the hyphae septate and thin-walled. Stipe cortex with rosettes.
Stipe cuticle of vertical to interwoven hyphae loosely arranged at the
surface and with somewhat elongate versiform free hyphal tips as caulocystidia, the hyphae refractive in KOH, with widely scattered dextrinoid particles present, no indications of slime either in KOH or in Melzer's.
Habit, habitat, and distribution. —On soil, in mixed woods, H. E.
Bigelow 12432 (TENN 35341), type, South Conway, New Hampshire,
September 4, 1963.
Observations. —The hoary, dark, viscid pileus, mild white latex, cinereous hoary stipe, broad lamellae, slightly colored spore deposit, and
nonstaining lamellae characterize this species. Sections of the stipe indicate that the surface is more likely not to be viscid than as Bigelow described it (viscid) in his notes.
139. Lactarius paludinellus Peck

Ann. Rep. N.Y. State Mus. 38: 133. 1885