The boletes of Michigan, by Alexander H. Smith and Harry D. Thiers.

156

THE BOLETES OF MICHIGAN

fuscous (no intermediate truly reddish state); FeSO4 staining flesh olivaceous.
Stipe 13 cm long, 2 cm thick at apex, 3.5 cm at base, whitish
overall, when cut soon "dusky brown" (violaceous-brown), lacking a red
stage in the color change, surface whitish and conspicuously roughened
by whitish points or squamules to almost reticulate in places, the ornamentation slowly brown to blackish on aging (no green, yellow, or blue
stains developing).
Spores (12) 13-15 X 4-5 u, subfusiform in face view, rather
obscurely inequilateral in profile, suprahilar depression shallow; smooth;
wall thickened somewhat, brownish ochraceous in KOH, in Melzer's yellowish or a few reddish tawny, no fleeting-amyloid reaction present.
Basidia 4-spored, 20-27 X 9-11.5 u, some with dark bister content
in KOH. Pleurocystidia rare, similar to cheilocystidia. Cheilocystidia
fusoid-ventricose, 26-35 X 8-12 u, apex obtuse, content either colored
or hyaline in KOH. Caulocystidia mostly clavate, 36-50 (70) X 10-20 i,
content smoky yellow in KOH, thin-walled, smooth, in fascicles. Caulobasidia rare.
Cutis of pileus a tangled layer of thin-walled, smooth, hyaline
hyphae 4-7 Mt wide, the terminal cells equal in diameter throughout their
length and obtuse at apex or varying to only slightly cystidioid, content
hyaline to pale dingy ochraceous in KOH, in Melzer's the content homogeneous (no sign of pigment globules), lacking truly inflated cells in the
cutis hyphae. Context of hyphae having red to orange-red content in
Melzer's in those near the cutis. Clamp connections none.
Habit, habitat, and distribution.-Under birch and aspen, Wildwood, Emmet County, July 14, 1963, Sherman Brough (Smith 66914).
Observations.-As dried the cutis of the pileus is somewhat cottony and so also is the ornamentation of the stipe. The pale color was
not an accident of development (such as the pileus being covered by
leaves). The species is closely related to L. clavatum, but differs in the
tree-associate, slightly smaller spores, different stature and in the hyphae
of the context next to the cutis being orange-red in Melzer's.
76. Leccinum subtestaceum Smith, Thiers, & Watling
Mich. Bot. 5:145. 1966
var. subtestaceum
Illus. Pls. 73-74.
Pileus 4-12 cm broad, convex to broadly convex, surface dark
ferruginous to liver color ("Hays-russet") and glabrous to finely fibrillose,