The boletes of Michigan, by Alexander H. Smith and Harry D. Thiers.

TYLOPIL US

123

Observations.-This, in our estimation, is clearly a Tylopilus even
though the color of the spore deposit remains to be recorded. It has the
aspect of T. rubrobrunneus, but is readily distinct because of its small
spores, mild taste, conspicuous cheilocystidia, and turf of pileocystidia
as a cuticle over the pileus. T. badiceps is one of the most easily recognized species in the genus.
55. Tylopilus sphagnorum (Peck) Smith & Thiers, stat. & comb. nov.
Boletus chrysenteron var. sphagnorum Peck, Bull. N. Y. State Mus. 150:64. 1910.
Illus. Figs. 50-51, 53.
"Pileus hemispheric or very convex, reddish-brown, the extreme
margin thin, slightly surpassing the hymenium, incurved, flesh white or
whitish; tubes longer than thickness of flesh.
"Pileus 2-3 cm broad; stem 2-4 cm long, 5-8 mm thick.
"Among sphagnum. Stow, Massachusetts, September. S. Davis."
The following data were obtained from the type: Spores 11-14 X
4-5 p, smooth, lacking an apical pore, shape in face view narrowly
elliptic to suboblong, in profile suboblong to somewhat inequilateral,
suprahilar depression usually broad and shallow, color in KOH yellowish
hyaline, in Melzer's yellowish to pale tawny, wall about 0.2 p thick.
Basidia 4-spored, 8-10 broad, hyaline in KOH and yellowish in
Melzer's. Pleurocystidia scattered, 28-42 X 8-14u, clavate to clavatemucronate or with a narrow short neck ending in a subacute apex, walls
thin, smooth, and hyaline, content homogeneous and ochraceous in
KOH, brown in Melzer's but soon fading to ochraceous, amorphous.
Cheilocystidia smaller than pleurocystidia but with the same yellow content.
Pileus cutis a tangled layer of broad (8-15 p) hyphae, the cells
mostly somewhat inflated, the end-cells voluminous (50-90 X 10-25 P),
clavate to fusoid-ventricose, thin-walled and smooth, content hyaline to
yellowish in either KOH or Melzer's, the cells not disarticulating and
typically are elongate. Hyphae of subcutis 4-10 p wide and with ochraceous content in Melzer's. Throughout the mounts of the cuticle were
found numerous patches of "amyloid" debris. Clamp connections absent.
Observations.-There can be little doubt that this fungus is a
Tylopilus. The gigantic end-cells of the cuticular hyphae indicate it is an
autonomous species. No pigment globules formed in Melzer's mounts of
the cuticular hyphae.