The boletes of Michigan, by Alexander H. Smith and Harry D. Thiers.

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THE BOLETES OF MICHIGAN

54. Tylopilus badiceps (Peck) Smith & Thiers, comb. nov.
Boletus badiceps Peck, Torrey Club Bull. 27:18. 1900.
Illus. Figs. 52, 54-55.
Pileus 4-8 cm broad, firm, convex to somewhat centrally depressed
when mature, dry, velvety, obliquely truncate on the margin, bay-red or
dark maroon. Context white, unchangeable but when bruised brown
spots slowly developing on standing, taste and color mild, sweet, suggestive of molasses. Tubes plane, adnate, white or whitish, becoming
dingy with age, the mouths minute. Stipe 4-5 cm long, 1.5-3 cm thick,
equal or slightly swollen in the middle, radiating, glabrous, solid, brownish.
The above description is adapted from McIlvaine's (1900) account.
He sent the specimens to Peck who described the species. Such a collection made by McIlvaine was found in the Peck collections by Stanley J.
Smith. It is without question the holotype. The following microscopic
data were taken from it.
Spores 8-10 X 3.5-4.5 I, smooth, lacking an apical pore; shape in
face view narrowly elliptic, in profile subelliptic or a weak suprahilar
depression causing the ventral line of the spore to be straight to slightly
concave; color in KOH hyaline to yellowish, in Melzer's yellowish, walls
0.2 p. thick.
Basidia 4-spored, 20-26 X 6-8 pu, hyaline in KOH, clavate. Pleurocystidia abundant, 27-38 X 8-12 j, narrowly ventricose-mucronate to
clavate-mucronate or more rarely fusoid-ventricose, content hyaline in
KOH, when well revived in Melzer's only weakly yellowish and homogeneous. Cheilocystidia abundant and prominent, similar to pleurocystidia but usually larger and walls distinctly yellow in KOH-possibly
from content adhering along the interior surface (the bright color present in poorly revived masses of cystidia). Caulocystidia not studied.
Tube trama of hyphae 4-8 I wide, tubular, hyaline, thin-walled,
transverse septa highly refractive and no amyloid reactions evident (the
tissue weakly yellow), hyphal arrangement of the Boletus subtype.
Pileus cuticle a compact palisade of pileocystidia 30-40 X 7-10 u,
obtusely fusoid-ventricose varying toward utriform or subclavatemucronate to clavate, the content yellow in KOH but this soon dissolving into the mount, in Melzer's dull reddish orange fading slowly to
dingy ochraceous, walls thin and with some debris adhering but no
characteristic pigmentation or wall roughening showing. Hyphae beneath
the palisade yellowish in Melzer's and laticiferous elements 4-7 u in
diameter with yellow content (in KOH or Melzer's) fairly numerous.
Clamp connections absent.