North American species of Mycena.

SUBDIVISIONS OF MYCENA

FOUR subgenera in Mycena-Mycenella, Pseudomycena, Glutinipes, and Eumycena-have been recognized. The traditional
concept of Mycenella has been enlarged here to include one section
with species having falsely echinulate spores, and the other with
those having nodulose spores. If these sections were to be recognized
as genera, they would then be Mycenella, characterized by species
with nodulose spores, and Fayodia, distinguished by the falsely
echinulate type of spore. The species of the latter have their closest
relationships in the Omphaliariae of Eumycena, and those of the
former in the Typicae. Pseudomycena has been used to include the
species of the section Basipedes of Fries (1872) characterized by the
presence of a basal disc. Glutinipes is defined to include all species
of both Mycena and Omphalia characterized by viscid stipes. Earle
(1909) gave the generic name Collopus to the viscid-stiped species of
Mycena. Ktihner (1938) has put these in various groups under his
section Mycenopsis of Eumycena, an arrangement which, in my
estimation, does not place sufficient emphasis on the viscidity of the
stipe. The series of more or less cespitose lignicolous forms in Glutinipes is related to members of the Typicae through M. laevigata.
The gray terrestrial species are related to the members of either the
Omphaliariae or the Typicae-M. quinaultensis and M. tenax to the
latter and M. clavicularis, M. odorifera, and M. vulgaris to the former.
The relationships of those grouped in the Viscosae are not so evident
as in those mentioned above, but they appear to be with the members
of Eumycena, possibly in the Calodontes. If one were to divide
Mycena into several genera, a procedure I do not now believe to be
desirable, Glutinipes would be one of the logical divisions.
Kiihner (1938) distinguished two subgenera, Eumycena for species
with amyloid spores and Paramycena for those with nonamyloid
spores. Singer, in 1938, also proposed an arrangement in which
several genera were recognized. Mycena was limited to species with
amyloid spores; the rough-spored species (they have nonamyloid
spores) were placed in Mycenella; and those remaining were divided
between Hemimycena and Pseudohiatula. My arrangement differs
sharply from the two mentioned above in putting secondary emphasis
on the amyloid reaction of the spores. Because of the difference in the
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