The North American species of Pholiota, by Alexander H. Smith and L. R. Hesler.

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The North American Species of Pholiota

HABIT, HABITAT, AND DISTRIBUTION: Scattered on hardwood logs,
Lewiston, Michigan, July 31, 1963, Smith 66988 (type), 66992.
OBSERVATIONS: This variety is distinguished primarily by the heavy
veil for the size of the basidiocarps.
17. Pholiota subechinata sp. nov.
Pileus 10-25 mm latus, late convexus, siccus, fibrilloso-squamulosus,
fulvus. Lamellae latae, subdistantes, adnatae, luteae demum fulvae. Stipes
2.5-3.5 cm longus, 1.5 mm crassus, flexilis, subluteus, deorsum fulvus.
Sporae 7-9 x 4-4.5 ii. Pleurocystidia desunt. Cheilocystidia 26-40(50) x
8-16 a., subclavata. Specimen typicum in Herb. Univ. Mich. conservatum
est; legit prope Mt. Adams, Wash. 19 Oct. 1954, Smith 49291.
Pileus 10-25 mm broad, convex to convex-depressed, surface dry and
minutely fibrillose-squamulose, squamules erect over disc and more
appressed near the margin, tawny to darker rusty brown. Context yellowish, thin, fairly lax, odor none, taste not recorded.
Lamellae broad, subdistant, adnate, seceding, dull yellow young,
darker fulvous than pileus when mature, edges flocculose.
Stipe 2.5-3.5 cm long, 1.5 mm thick, equal, fairly pliant, dull yellowish above, soon dark fulvous from the base up (over all as dried), thinly
coated to the evanescent veil-line from the remnants of the veil, with a
pad of yellowish mycelium at base.
Spores 7-9 x 4-4.5,u, smooth, germ pore very minute, dingy ochraceous in Melzer's reagent, in KOH cinnamon, oblong to elliptic in face
view and some obscurely angular, somewhat to distinctly bean-shaped in
profile.
Basidia 4-spored, 22-30 X 5-7,u, yellowish hyaline in KOH, narrowly
clavate. Pleturocystidia none. Cheilocystidia 26-40 (50) X 8-16 pF, mostly
somewhat enlarged at apex (the widest part) but apex varying from
rounded to truncate or subacute, rarely bifurcate, walls thin, smooth and
hyaline in KOH (reminding one of those of P. crinaceella but not as
capitate). Caulocystidia with rusty cinnamon thin walls usually smooth,
35-60 X 8-12 /A, fusoid to boomerang-shaped, terminal or arising laterally from hyphae of stipe cortex.
Gill trama regular, of yellowish to brownish hyphae with smooth
scarcely thickened walls; subhymenium not distinctive. Pileus cutis a
trichodermium of hyphae mostly with short broad cells (1-2 times as long
as broad), the walls slightly thickened or thin and many with coarse
plates of incrusting material, in KOH dark red (near "Bay"), end-cells
subglobose to elliptic or ovate. Clamp connections present. All tissues
inamyloid.
HABIT, HABITAT, AND DISTRIBUTION: On a conifer log, on the upper
Cispus River, Washington, near Mt. Adams, Oct. 19, 1954, Smith 49291
(type).
OBSERVATIONS: The sphaerocyst-like cells in the trichodermial hyphae