The North American species of Pholiota, by Alexander H. Smith and L. R. Hesler.

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The North American Species of Pholiota

color change in drying and the variation in shape of the pleurocystidia
make it amply distinct.
195. Pholiota subflavida (Mut1.) comb. nov.
Gymnopilus subflavidus Murrill, 4: 252. 1912.
Flamnmula subflavida Murrill, Mycologia 4: 262. 1912.
Illustrations: Text figs. 436; 440-441; pl. 88.
Pileus 3-5 cm broad, thin, conic or convex to expanded, umbonate
when young, melleous with fulvous center, becoming green-spotted when
handled, slimy, glabrous, smooth, margin entire, strongly incurved.
Lamellae sinuate or adnate, citrinous to fulvous, medium broad,
subdistant.
Stipe 4-7 cm long, 5-8 mm thick, equal, cremeous above, pale
fulvous below, smooth, fibrillose. Veil slight, citrinous, membranous in
young stages, soon breaking into fibrils and leaving no annulus.
Spores 5.5-7.5 (8.5) X 3.5-4.5 (5) p,, smooth, apical pore small but
distinct; shape in face view elliptic to ovate, in profile obscurely inequilateral; color tawny in KOH under microscope, in Melzer's reagent about
the same color; wall about 0.25 C/ thick.
Basidia 17-22 x 4-5 bt, 4-spored, clavate, yellowisl and KOH and
Melzer's reagent. Pleurocystidia abundant, 48-72 x 9-15 p-, fusoid-ventricose, apex subacute to obtuse, walls thin and smooth, content ochraceous
tawny to yellowish fading to hyaline and homogeneous. Cheilocystidia
28-45 (60) x 8-13,u, subfusoid with obtuse apex varying to fusoid-ventricose, walls thin, some with debris adhering, content homogeneous. Caulocystidia not studied.
Gill trama a central area of floccose hyphae, the cells yellowish in
KOH (poorly revived); subhymenium of narrow gelatinous hyphae in
a distinct layer. Pileus cutis a thick gelatinous layer of hyaline to yellowish hyphae 2-4,L diam.; hypodermial region of bright fulvous hyphae
with incrusted walls, cells 4-12 /x diam. Context hyphae interwoven,
smooth, yellowish in KOH. Clamp connections present. All hyphae inamyloid. (Above microscopic data all from type).
HABIT, HABITAT, AND DISTRIBUTION: On conifer debris or near it, fall,
Idaho and WNashington. Type studied.
OBSERVATIONS: This is another member of the P. spumnosa complex
which has been named as a distinct species from North America. We
suspect that the spotting green when handled is some sort of artifact due
to wet weatler and the general tendency of olive tints to develop in the
context in age. The type specimen reminds one of P. spunmosa. We recognized the species here as an aid to future studies, but Smith in all of his
collecting in the Northwest has not been able to recognize it on any other
feature than the smaller spores. It may well be merely a small spored
variety of P. spumosa. P. subfulva has thick-walled pleurocystidia.
We have a number of collections from Idaho (Smith 73387, 73388,