The North American species of Pholiota, by Alexander H. Smith and L. R. Hesler.

Smith * Hesler

343

thin-walled, wall hyaline to slightly yellowish; content homogeneous or
with a plug of yellowish colloidal material (in KOH) in apical part,
refractive particles present in this material in some, or present in cells
not showing the colloidal material. Cheilocystidia 30-45 x 12-20 /U
broadly subutriform to clavate, thin-walled, smooth, walls yellowish to
hyaline, content yellowish to hyaline and homogeneous. Caulocystidia
none found.
Gill trama a central strand of floccose thin-walled, yellowish parallel
to somewhat interwoven hyphae 4-10 /u diam. or cells finally more inflated; subhymenium a thick layer of narrow gelatinous hyaline hyphae.
Pileus with a thick gelatinous pellicle of narrow (2-5 u) hyaline to
yellowish smooth to incrusted hyphae grading into a hypodermial region
of heavily incrusted brown hyphae 4-12 /t diam. Context of yellowish
smooth hyphae, thin-walled (0.25-0.3 /u estimated), interwoven, and
with cells inflated to 15 /x. All hyphae inamyloid and hyphae at apex of
stipe yellowish to hyaline in KOH. Clamp connections present.
HABIT, HABITAT, AND DISTRIBUTION: Among mosses and Calamagrostis arundinacea on bouldery slope under beech and birch. Sweden (Lundell 5683).
OBSERVATIONS: That the problem of the identity of P. lubrica is an
acute one is an understatement. Konrad (1936) gave the spore size as
8-9.5 x 4.5-5.5,u. For this reason our data are all taken from a Lundell
(no. 5683) specimen collected in Sweden. A Moser specimen studied by
Hesler had thick walled cystidia and Moser (1953) gives the spores as
5-6 x 3-4 pi, a broader spore in relation to its length than we have
found in any of this group. Fries' illustration Icones pl. 116 of Agaricus
(Flammula) lubricus shows an ochraceous marginal area to the pileus
and a brown disc. The stipe is white, staining brown at the base. In
Systema, however, Fries described the pileus as "cinnamomeus." This is
in line with (copied from) Persoon's description and hence is accepted
as authentic for the color.
The following is a description of collections Smith 73892, 73678,
74181, and 74071 all from Upper Priest Lake in Idaho:
Pileus 4-8.5 cm broad, broadly convex with an incurved margin,
expanding to plane or shallowly depressed, surface with patches of the
pallid fibrillose veil over marginal area on the slime covering the surface,
slime often dripping from edge of pileus; color reddish tawny over disc
and pallid over margin mature, when very young the margin ivory
yellow; veil pallid, leaving a fringe on margin at first. Context whitish
with watery brownish line next to gills; taste mild, odor none, with
FeSO4 green.
Lamellae whitish becoming dull cinnamon, broad, bluntly adnate,
crowded, edges even.
Stipe 6-8 cm long, 9-11 mm thick, equal, pallid, soon rusty red to
bay below, whitish above, fibrillose scurfy from remains of the veil, apex
silky and yellowish pallid.