The North American species of Pholiota, by Alexander H. Smith and L. R. Hesler.

Smith ~ Hesler

279

5. Stipe 1.5-4 (5) mm thick; veil pallid at first.........P. highlandensis
6. Pileus pale buff young; veil orange brown on aging..... P. molesta
6. Pileus dark yellow-brown; veil lemon yellow young.............................. P. brunnescen s
148. Pholiota carbonaria A. H. Smith, Mycologia 36: 253. 1944.
Pholiota carbonicola Singer, Agar. Mod. Tax. p. 543. 1963.
Illustrations: Text figs. 322-324; pl. 68.
Pileus 2-4 cm broad, convex with an incurved margin beautifully
appendiculate with remains of the veil, expanding to plane or with slight
umbo, rarely the disc slightly depressed, "warm-buff" to "antimonyyellow" young, becoming darker and finally nearly cinnamon-brown in
age on the disc, the margin remaining paler, surface viscid, covered at
first by concentric rows of small nearly "hazel" or "ferruginous" fibrillose
scales, more or less glabrescent. Context watery brown, thick in the disc;
odor and taste not distinctive.
Lamellae bluntly adnate, narrow, crowded, equal, white to grayish
when young, becoming nearly snuff-brown at maturity, edges slightly
crenulate.
Stipe 3-6 cm long, 4-6 mm thick, equal, somewhat compressed or
terete, fibrous, dingy watery yellowish within, solid, becoming tubular
at least in apical region, surface covered by small "hazel" to "ferruginous"
fibrillose squamules from the remains of the veil, squamules often recurved, ground yellowish but darkening slightly below, apex pruinose.
Spores (5) 6-7 (8) x 3.5-4 (4.5) a, smooth, apical pore present but
apex not truncate; shape in face view elliptic to ovate, in profile subelliptic to obscurely inequilateral; color tawny in H-,O mounts fresh,
revived in KOH cinnamon-brown, in Melzer's reagent paler cinnamon;
wall about 0.3 I/ thick.
Basidia 18-23 x 6-7 JL, 4-spored, clavate, yellowish to hyaline in
KOH or Melzer's reagent. Pleurocystidia abundant, 50-88 x 9-14 /u,
fusoid ventricose with obtuse or rarely subcapitate apices, when fresh
some with adhering amorphous material around the apex, smooth in
KOH, hyaline or with yellowish content in the neck or throughout, thinwalled, rarely the apex forked (in Smith 54261); cheilocystidia abundant,
25-47 x 7-13 /u, clavate to subfusoid or obscurely fusoid ventricose, walls
ochraceous in KOH, smooth or some slightly incrusted. Caulocystidia
none.
Gill trama hyaline, interwoven, floccose, some sections showing a
slight gelatinization of the subhymenium. Pileus trama floccose-interwoven and yellowish hyaline in KOH. Hypodermium rusty brown.
Cuticle gelatinous, hyaline to yellowish, of interwoven hyphae 2-3.5 u
in diameter. Clamp connections present. All hyphae inamyloid.
HABIT, HABITAT, AND DISTRIBUTION: On burned soil and exposed