The North American species of Pholiota, by Alexander H. Smith and L. R. Hesler.

Smith * Hesler

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along the edges when rubbed, close to crowded, 3 tiers of lamellulae,
moderately broad to broad in age, edges even.
Stipe (3)5-10(12) cm long, (3)5-10 mm thick, equal or slightly
enlarged at base, solid, stuffed to tubular, concolorous with pileus inside
and out, darkening to cadmium yellow at base, covered up to the annulus
with a dense coating of recurved yellow scales from the ruptured veil,
silky fibrillose above the annulus. Veil forming an evanescent, superior
zone of bright yellow fibrils or submembranous.
Spores 4-5 x 2.5-3.t smooth, ochraceous in KOH and about the
same color in Melzer's reagent, oblong to ellipsoid, smooth, wall thin,
apical pore not evident.
Basidia 18-22 x 3.5-4.5 pt, narrowly clavate, 4-spored, hyaline in
KOH, yellowish in Melzer's reagent. Pleurocystidia of two types; 1) 25 -35 x 6-8 pf, clavate to mucronate and with a dark flavous homogeneous
content as revived in KOH, in Melzer's reagent yellowish, walls smooth
thin and hyaline; 2) clrysocystidia 26-40 x 6-9 pJt fusoid ventricose to
clavate-mucronate, content amorphous and granular-aggregated to forming folded (reticulate) masses cinnalbar-color to orange-brown in Melzer's
reagent. (Cheilocystidia as in type 1 of pleurocystidia or small and
cylindric (18-20 x 3 pt), the former abutndant. Caulocystidia present
only as filamentosc end-cells of surface hyphae.
Gill trama of floccose subparallel hypha 4-8 pt 1)road, ocllraceous
in KOH and pigment diffusing into nmount, masses of needle-like crystals
forming in the mount; cell walls thin, smootl, yellowisll; sbl)lhymenium
of narrow hyaline somewhat interwoven gelatinous hyphae. Pileus cutis
a thick layer (100 pt +) of narrow (2-4 pu) gelatinous hyphae yellowish
in KOH; hypodermial region of oclraceous-brown hyphae, structurally
continuous with context, the hyphae compactly interwoven, 4-12 p. diam.,
walls smooth thin and oclraceous to hyaline; oleiferous hyphae present.
Clamp connections present. All hyphae inamyloid.
HABIT, HABITAT, AND DISTRIBUTION: On conifer logs and stumps,
Maine, Michigan, Tennessee, North Carolina, Idlaho, an(d Washington;
August-October.
OBSERVATIONS: Kauffman, when collecting in the Olympic Mountains of Washington in 1925 described viscid specimens as a tentative
new species. When Smith was working in the same area he found the
same variation in the same condition and since a viscid as contrasted to
a non-viscid pileus was an important feature in Pholiota he described the
viscid variant naming it in honor of Kauffman. However, anatomical
studies of this variant in comparison with material collected by Kauffman in Sweden and presumably checked by Romell, and material sent
to Smith from France by Josserand are all so much alike microscopically
that there appears to be no clear line of separation-particularly as
applied to the pileus cutis. In addition, during the last twenty years
Smith has observed the species in many areas of the United States and
under variable weather conditions. He is now convinced that the prol