The North American species of Pholiota, by Alexander H. Smith and L. R. Hesler.

Smith ~ Hesler

189

buff or darker where bruised, almost equal, slightly tapering upward,
solid and whitish within. Annulus membranous, persistent, 3-4 mm
broad, median, deflexed, colored by the spores.
Spores 6-8.5 x 3.7-4.5 A/, elliptic to subovate, in profile slightly
inequilateral; smooth, germ-pore minute.
Basidia 23-38 X 9-15 ut, 4-spored. Pleurocystidia of two kinds: 1)
chrysocystidia 25-28 x 9-15 /,, clavate-mucronate to subfusoid, often with
an amorphous content; 2) leptocystidia 50-54 X 10-17 p,, fusoid-ventricose to subcylindric, erratic in occurrence. Cheilocystidia 45-54 x 13-16,u, fusoid-ventricose.
Gill trama slightly interwoven, hyphae 3-6 /a broad; subhymenium
not distinctive. Pileus trama radial, somewhat interwoven. Cuticle not
sharply differentiated, not viscid, the surface bearing some fascicles (the
scales) of brown hyphae 6-12 u/ broad and usually faintly incrusted.
Clamp connections present.
HABIT, HABITAT, AND DISTRIBUTION: On soil, solitary, in dense coniferous forests, Oregon, October. Type studied.
OBSERVATIONS: We include this species with some reservations. A
shiny pileus is not consistent with the scaly condition as described. It
seems amply distinct as a species on macroscopic features, and the spores
are certainly Pholiota-like.
Stirps Squarrosa
Key
1. Spores 6-8 /L long; some pleurocystidia hyaline in KOH and evenly
refractive throughout........................................ P. squarrosa
1. Spores smaller (5-6.5 / long); pleurocystidia not as above........................................................................................................... P. k o d i a k e n s i s
96. Pholiota squarrosa (Fr.) Kummer, Der Fiihrer in die Pilzkunde.
p. 84. 1871.
Agaricus squarrosus Fries, Syst. Myc. 1: 243. 1821.
Dryophila squarrosa (Fr.) Quelet, Enchir, Fung., p. 68. 1886.
Hypodendrum floccosum (Schaeff.) Overholts, North Amer. Fl. 10: 280.
1932.
Var. squarrosa
Illustrations: Pls. 41-43.
Pileus, 3-10 (12) cm broad, obtuse to convex when young, the margin incurved, becoming campanulate with a conic umbo to broadly
convex or nearly plane, surface dry, cuticle broken up into numerous
innate, recurved, or squarrose, scales, the margin at first conspicuously
fringed with veil remnants, cuticle sometimes denuded along the edge