The North American species of Pholiota, by Alexander H. Smith and L. R. Hesler.

Smith ~ Hesler

179

homogeneous. Caulocystidia 33-60 x 6-9 Iu, cylindric to narrowly clavate,
hyaline in KOH, thin-walled, content "empty."
Gill trama subparallel, of floccose-interwoven thin-walled hyphae;
walls yellowish in KOH, smooth, content often orange-brown in Melzer's
reagent. Subhymenium not a sharply defined layer, hyphae compactly
interwoven, non-gelatinous (in sections the layer appearing more or less
cellular). Pileus cutis a gelatinous pellicle of mostly ochraceous narrow
(2-3 Iu) gelatinous hyphae smooth to incrusted, hypodermial region of
ochraceous, incrusted floccose hyphae 4-10 pX diam. Context hyphae
interwoven, filled with "colloidal" content, becoming rusty reddish in
Melzer's reagent. Clamp connections present.
90a. Pholiota flavida var. graveolens var. nov.
Illustrations: P1. 35.
Pileus 5-7 cm? lants, late convexius, visciduis, glaber, sordide lnitens.
Contextus liteolus; odor graveolens. Lamellae adnatae, latae confertae,
liteae demrum castaneae. Stipes 9-13 cm longus, 7-9 mm crassus, sursum
hlteolius, deorsumn subcastanens, sparse fibrillosus. Sporae 8-10 x 4.5-5 p.
Pleurocystidia desnlt. Cheilocystidia 23-36 X 5-8 pu, subventricosis sacpe
irregularis. Specimen typicum in Herb. Univ. Mich. conservatum est; legit
prope Priest Lake, Idaho. 26 Sept. 1964. Smith 70814.
Pileus 5-7 cm broad, plane with a decurved margin, viscid, glabrous,
dingy honey-tan where not covered with spores, fading to dingy ochraceous (chestnut where covered with deposited spores in age). Context
pliant; odor heavy and like some fats, taste mild; FeSO4 no reaction, KOH
no reaction.
Lamellae adnate-seceding, dingy yellow, becoming "auburn" from
spores, horizontal, fairly broad, close, edges even.
Stipe 9-13 cm long, 7-9 mm thick, equal, yellowish pallid at apex,
dark rusty chestnut from base up, thinly fibrillose from a very thin veil
which leaves a few indistinct patches on the cap margin, cortex colored.
Spore deposit "chestnut" to "auburn" (with a strong red tinge).
Spores 8-10 x 4.5-5 p,, smooth, apical pore distinct, wall thickened
(0.3-0.4,u); shape narrowly elliptic to ovate in face view, in profile
somewhat inequilateral to subovate; dingy cinnamon or paler in KOH,
more reddish cinnamon in Melzer's reagent.
Basidia 23-29 x 7-8 iu, 4-spored, yellowish in KOH or Melzer's.
Pleurocystidia none; cheilocystidia 34-47 X 4-12 pi ventricose near base
and with a crooked neck ending in an obtuse apex. Thin-walled. Gill
trama with an indistinct cellular hymenium, central area of smooth inflated hyphae (to 25 u). Pileus epicutis a pellicle of gelatinous hyphae
2.5-6 ip diam., yellow in KOH but fading; hypodermium yellow in KOH.
Clamp connections present.
HABIT, HABITAT, AND DISTRIBUTION: Caespitose at base of a conifer
stub, near Priest Lake, Idaho, Sept. 26, 1964. Smith 70814.