The North American species of Pholiota, by Alexander H. Smith and L. R. Hesler.

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The North American Species of Pholiota

OBSERVATIONS: This is a very odd Pholiota because of the long rooting base often giving rise to a number of secondary stipes (merismoid)
and with adhering sand much as in Laccaria trullisata, the rather broadly
ellipsoid spores, and the relatively short pleurocystidia for the group.
It's relationships appear to be in the P. lubrica complex but the spores
at once distinguish it from all of these.
189. Pholiota foedata (Pk.) comb. nov.
Hebeloma foedatum Pk., Bull. Torrey Bot. Club 22: 202. 1895.
Gymnopilus foedatus (Pk.) Murrill, North Amer. Fl. 10: 206. 1917.
Cortinarius foedatus (Pk.) Kauffman, Amer. Journ. Bot. 13: 28. 1926.
Illustrations: Text figs. 425-426.
Pileus 3.5-7.5 cm broad, convex, becoming plane or centrally depressed, reddish cinnamon, very viscid or glutinous, glabrous.
Lamellae emarginate and with a decurrent tooth, cinnamon colored,
becoming mummy brown, medium broad, subcentricose, crowded.
Stipe 3.5-6.5 cm long, 4-8 mm thick, paler than the pileus, fibrillose,
equal or slightly thickened at the base, solid.
Spores 6-7.5 x 4-4.5,u, smooth, apical pore minute, wall up
to 0.25-0.5 /u thick, shape elliptic to oval in face view, elliptic to very
obscurely inequilateral in profile, in KOH rusty cinnamon and about
the same color in Melzer's reagent.
Basidia 18-24 x 5-6 u, 4-spored, clavate, yellowish-hyaline in KOH,
yellowish in Melzer's reagent. Pleurocystidia abundant, 36-70 x 9-15 /u
fusoid-ventricose, apex obtuse, wall thin and smooth in KOH but content
at times reticulate-wrinkled and ochraceous in KOH as coagulated (often
clearing on standing). Cheilocystidia similar to pleurocystidia or shorter
and more fusoid, apex obtuse. Caulocystidia none found on type or on
Sipe 248.
Gill trama a central strand of floccose hyphae not distinctively
colored as revived in KOH, the walls thin and smooth; subhymenium
a layer of hyaline branched gelatinous hyphae 2-3 jL diam. Pileus cutis
a gelatinous pellicle of yellowish collapsed hyphae as revived in KOH;
hypodermium of rusty brown hyphae 3-7 /p diam. with rusty colored
incrustations. Context hyphae smooth and thin-walled, yellowish in KOH,
hyphal cells greatly inflated. All hyphae inamyloid. Clamp connections
present.
HABIT, HABITAT, AND DISTRIBUTION: Along streets, or among oak leaves.
California (Pasadena). Type studied.
OBSERVATIONS: The type was from under oak in southern California
and had adnexed gills. McClatchie's specimens which we have studied
had dark reddish-rust colored spores much as in P. subangularis but in
Sipe's 248 from conifer duff in the mountains they were paler. The gill
color of McClatchie's material seen by us is now rusty cinnamon not a
blackish brown (mummy brown). For these reasons the microscopic