The North American species of Pholiota, by Alexander H. Smith and L. R. Hesler.

162

The North American Species of Pholiota

Stipe 4-7 cm long, 3-7 mm thick, flexuous, fibrillose, yellow, darkening toward the base, apex enlarged.
Spores (5.5)6-7.5 x 3.5-4 Ax, smooth, apical pore distinct but apex
not truncate; shape oblong in face view varying to elliptic, in profile
obscurely inequilateral to subelliptic or obscurely bean-shaped; in KOH
pale tawny and in Melzer's reagent paler.
Basidia 18-22 x 4.5-6 /u obese, 4-spored, yellowish to hyaline in
KOH, yellowish in Melzer's reagent and with numerous yellowish globules. Pleurocystidia scattered, imbedded, 28-37 X 9-12 bu subglobose,
ovate in optical section or elliptic, with a short pedicel; with granularreticulate ochraceous content in KOH, thin-walled, some debris adhering on wall, the content darker (yellow-brown) in Melzer's than
in KOH. Cheilocystidia versiform, 24-48 x 3-7 /u, contorted-filamentous,
filamentous-capitate, narrowly fusoid-ventricose, or any of these shapes
proliferating at apex (see figs.), walls thin to slightly thickened (-0.5 AL),
yellowish in KOH, smooth, content homogeneous. Caulocystidia resembling the cheilocystidia or greatly elongate-subcylindric with a slight
thickening of the wall at obtuse apex (50 x 5 /u ~), smooth, content
homogeneous.
Gill trama of subparallel floccose hyphae with greatly elongated
cells 5-12 / wide; walls thin, yellowish in KOH, smooth; subhymenium a
gelatinous zone of interwoven narrow (2-3 AL) hyaline hyphae. Pileus
cutis a gelatinous pellicle of narrow ochraceous hyphae; hypodermial
region of rusty to yellowish-tawny hyphae with smooth to slightly incrusted walls. Context hyphae interwoven, cells greatly inflated, walls
smooth and yellowish in KOH. Clamp connections present. All hyphae
inamyloid.
HABIT, HABITAT, AND DISTRIBUTION: Caespitose, on soil and near
buried wood, Oregon, October-November.
OBSERVATIONS: This is distinguished by its honey odor and its pileus
with an incurved margin. It is close to P. ornatula, which has smaller
cheilocystidia and subdistant, pallid gills. The pleurocystidia are not
typical chrysocystidia but because of their distinctive content would have
to be classified there in a broad sense of the term. Apparently the cheilocystidia may proliferate more than once as several were seen in which it
was thought that at least 3 stages could be seen.
MATERIAL EXAMINED: OREGON. Smith 68780, 68781 (type, from
Portland, collected by Ruth Oswald, November 2, 1957).
82. Pholiota flavescens sp. nov.
Illustrations: Text figs. 113-116.
Pileus 1-4 cm latus, convexus, albidus demum pallide luteus, vel
flavidus, saepe ad marginem luteo-olivaceus. Lamellae subdecurrentes,
pallide hIteae, confertae, latae. Stipes 2-4 cm longus, 2.5-4(8) mm cras